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Species Profile

Desert Tortoise

Gopherus agassizii

Built for burrows, made for deserts

Desert Tortoise Distribution

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Endemic Species
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At a Glance

Wild Species
Also Known As Agassiz's tortoise, tortuga del desierto
Diet Herbivore
Activity Diurnal+
Lifespan 60 years
Weight 7 lbs
Status Vulnerable
Did You Know?

Adults typically measure ~20-36 cm straight carapace length (max ~38 cm) and often weigh ~4-7 kg.

Scientific Classification

A North American land tortoise adapted to arid habitats, known for living in burrows, slow growth, long lifespan, and herbivorous diet dominated by desert annuals and perennials.

Kingdom
Animalia
Phylum
Chordata
Class
Reptilia
Order
Testudines
Family
Testudinidae
Genus
Gopherus
Species
Gopherus agassizii

Distinguishing Features

  • High-domed brownish carapace and sturdy, elephantine hind limbs typical of tortoises
  • Strong forelimbs for digging; spends substantial time in burrows
  • Herbivorous, relying on seasonal desert vegetation and using burrows to manage heat and water stress

Physical Measurements

Males and females differ in size

Height
5 in (4 in – 6 in)
Length
12 in (8 in – 1 ft 3 in)
Weight
11 lbs (8 lbs – 15 lbs)
10 lbs (7 lbs – 14 lbs)
Top Speed
0 mph
About 0.48 km/h; usually slower

Appearance

Primary Colors
Secondary Colors
Skin Type Thick, dry, heavily keratinized reptilian skin with large, pebbly scales; forelimbs robust with enlarged, spade-like scales for digging; horn-like scales present on limbs.
Distinctive Features
  • Adult Desert Tortoise (Gopherus agassizii) carapace usually 20–36 cm long, sometimes up to about 38 cm; body mass commonly 3–7 kg, larger adults may weigh more.
  • High-domed shell with strong forelimb digging adaptations: stout forelegs, flattened/spade-like forefeet, and enlarged scales used to excavate burrows (a key Mojave survival trait).
  • Prominent beak-like mouth (tomium) suited to herbivory; head and limbs are robust and scaled rather than smooth-skinned (terrestrial tortoise, not aquatic).
  • Gular scutes (front of plastron) can project forward; males often develop a more pronounced gular projection used in combat/ramming.
  • Agassiz's desert tortoise spends much of the year in burrows to avoid deadly heat and drying out. It stores water in its bladder and reuses it during droughts.
  • Burrow use is visually/field-relevant: individuals frequently show shell abrasion/scuffing and dust staining from repeated burrow entry/exit and basking at burrow mouths; burrows are used for winter brumation and summer heat avoidance.
  • Gopherus agassizii is the Mojave Desert tortoise, mostly west of the Colorado River. Gopherus morafkai is the Sonoran Desert tortoise east/southeast of the river. Genetic and body-shape studies support this split.
  • Mojave desert tortoise populations are listed as Threatened under the U.S. Endangered Species Act. Major threats: habitat loss and fragmentation, predators (especially ravens), upper respiratory disease (Mycoplasma), and roads/OHV.
  • Different from Gopherus morafkai (Sonoran desert tortoise) in range (separated by the Colorado River) and genetics. Shell and scale traits are subtle, so location and expert help are used; management and laws differ.

Sexual Dimorphism

Sexual dimorphism is moderate and most evident in plastron shape, gular projection, tail length, and chin gland development; males also tend to be larger on average in many populations.

  • Plastron typically concave (facilitates mounting during mating).
  • Gular projection (anterior plastron) generally longer/more pronounced; used in male-male combat/ramming.
  • Tail longer and thicker with vent positioned farther from shell edge than in females.
  • Chin glands larger/more active; males often show more pronounced courtship/territorial behaviors near burrows.
  • Plastron generally flat to slightly convex.
  • Gular projection shorter/less protruding than in males.
  • Tail shorter with vent closer to shell edge.
  • Body form may appear broader posteriorly when gravid; females allocate substantial resources to egg production in favorable rainfall years (seen in seasonal body condition changes).

Did You Know?

Adults typically measure ~20-36 cm straight carapace length (max ~38 cm) and often weigh ~4-7 kg.

They can spend ~95% of their lives in burrows or rock shelters, avoiding lethal surface heat and cold.

Females commonly lay 1 clutch/year (sometimes 2), with ~1-14 eggs per clutch (often ~4-6).

They grow slowly and often don't reach sexual maturity until ~15-20 years old.

A large urinary bladder lets them store water and reabsorb it-vital for surviving long dry periods.

They are a "keystone" burrow-maker: dozens of other desert animals use tortoise burrows for refuge.

This is the Mojave species: it lives mostly north/west of the Colorado River-unlike the Sonoran desert tortoise (Gopherus morafkai) to the south/east.

Unique Adaptations

  • Extreme water economy: large bladder storage plus physiological reabsorption helps endure drought; can capitalize quickly on rainfall by drinking from puddles/soil depressions.
  • Heat-avoidance engineering: burrows buffer temperatures and humidity, creating a stable microclimate compared with the desert surface.
  • Tough, keratinized forelimbs and strong claws specialized for digging in packed desert soils.
  • Salt/nitrogen management suited to arid diets: excretes uric acid (less water loss than urea) and tolerates dry forage periods.
  • Low metabolic rate and the ability to reduce activity for months (brumation/estivation-like dormancy) to survive resource scarcity.
  • Shell and skin reduce water loss and provide protection, but survival depends heavily on behavioral thermoregulation (timing and shelter use).

Interesting Behaviors

  • Burrow construction and maintenance: digs long, gently sloped tunnels used for thermoregulation, predator avoidance, and winter brumation.
  • Seasonal activity pulse: most surface activity occurs in spring (and some in fall); during extreme summer heat they retreat to burrows for long periods.
  • Basking and shuttling: alternates between sun and shade/burrow entrances to keep body temperature within a safe operating range.
  • Herbivorous foraging strategy: targets spring annual wildflowers and grasses when water/nutrients are highest; shifts to tougher perennials as the season dries.
  • Reproduction timing: courtship and mating peak in late summer-fall; nesting typically late spring-early summer with temperature-dependent incubation (~90-120+ days).
  • Site fidelity: many individuals repeatedly use familiar burrows, shelter sites, and movement corridors within a home range.
  • Defense behaviors: pulls head/limbs into shell, uses forelimbs as "shields," and may wedge into burrow entrances against predators.

Cultural Significance

Agassiz's desert tortoise (Gopherus agassizii) is a key Mojave species, listed Threatened under the U.S. Endangered Species Act and CA/NV state reptile. Its burrows shelter many species. Threats: habitat loss, Mycoplasma, wildfire, invasive grasses, ravens, and climate change. Found northwest of the Colorado River, unlike Sonoran tortoise (G. morafkai).

Myths & Legends

"Turtle Island" creation traditions (widely told among many Indigenous peoples of North America) describe the world formed on the back of a turtle/tortoise-an enduring cultural symbol of stability, endurance, and life-bearing earth.

In Southwestern Indigenous storytelling, tortoise is often portrayed as a patient, resilient figure whose survival comes from persistence and wise use of shelter and water-themes that echo the desert tortoise's burrow-centered life.

The species name agassizii honors naturalist Louis Agassiz. Early 1800s scientific descriptions helped make the Desert Tortoise (Gopherus agassizii) a symbol of the American desert in museums and books.

Conservation Status

VU Vulnerable

Facing a high risk of extinction in the wild.

Population Decreasing

Protected Under

  • United States Endangered Species Act (ESA): Mojave population listed as Threatened (listed 1990; critical habitat designated in subsequent rulemakings).
  • CITES Appendix II (international trade regulated for Gopherus spp., used to limit commercial exploitation).
  • State protections in core range states (e.g., California and Nevada prohibit unauthorized take/possession; regulations vary by jurisdiction).
  • Large areas occur within protected lands and conservation designations (e.g., Desert Tortoise Conservation Areas, ACECs, National Parks/Monuments), with management focused on limiting surface disturbance, reducing road mortality, and controlling subsidized predation (USFWS/BLM frameworks).

Life Cycle

Birth 6 hatchlings
Lifespan 60 years

Lifespan

In the Wild
50–80 years
In Captivity
60–100 years

Reproduction

Mating System Polygynandry
Social Structure Solitary
Breeding Pattern Transient
Fertilization Internal Fertilization
Birth Type Internal_fertilization

Agassiz's Desert Tortoise (Gopherus agassizii) shows a non‑territorial, polygynandrous system: solitary adults mate with multiple partners during brief spring and late‑summer/fall encounters. Males search and court; females store sperm, lay 1–14 eggs, and provide no care.

Behavior & Ecology

Social Aggregation Group: 1
Activity Diurnal, Crepuscular
Diet Herbivore Moist, nutrient-rich spring annual forbs-especially desert plantain (Plantago ovata) and desert dandelion (Malacothrix glabrata) when abundant after winter rains.
Seasonal Hibernates

Temperament

Generally calm and non-aggressive outside the breeding season; avoids conflict by retreating to burrows.
Seasonal male-male aggression and dominance interactions during breeding: head-bobbing, circling, gular-horn probing, shell ramming, and occasional biting; males may attempt to overturn rivals (documented in Gopherus courtship/agonistic behavior; e.g., Woodbury & Hardy 1948; USFWS 2011).
Risk-avoidant/cryptic: strong tendency to use burrows for thermoregulation and predator avoidance; juveniles especially secretive and spend proportionally more time sheltered (USFWS 2011).

Communication

Hissing/expulsive sounds associated with rapid air movement during retraction or stress handling Common in tortoises; reported for Gopherus in field handling contexts
Low grunts/exhalation sounds can occur during close-contact courtship/mating Not a primary signaling mode; described anecdotally and in behavioral accounts for Gopherus spp.
Visual signaling: stereotyped head-bobbing and postural displays in courtship and male-male interactions; approach/parallel walking and circling prior to mounting attempts Woodbury & Hardy 1948; USFWS 2011
Tactile signaling: shell ramming, pushing, gular-horn contact/probing, and mounting behaviors; physical contests can determine access to mates or burrows in a local area.
Chemical/olfactory cues: scent investigation Sniffing) of conspecifics and substrates; presumed use of glandular secretions (chin/gular region) and cloacal cues in mate assessment and individual recognition (reported broadly for Gopherus social behavior; USFWS 2011
Spatial/indirect social information "HUBS" pattern): shared use of burrow networks and repeatedly used burrows creates contact hubs where individuals encounter scent trails and signs; this produces population-level connectivity despite solitary living, with variation by sex (males typically range more widely) and by rainfall/forage availability (USFWS 2011; Berry & Christopher 2001

Habitat

Biomes:
Desert Hot
Terrain:
Plains Valley Hilly Rocky Sandy
Elevation: -3386 in – 5249 ft 4 in

Ecological Role

Large, long-lived desert herbivore and ecosystem engineer that links rainfall-driven primary production to higher trophic levels and modifies habitat via burrowing.

Seed dispersal via endozoochory for some consumed plants (seeds can pass through the gut and be deposited with nutrient-rich feces in microsites) Herbivory that can influence plant community composition and seasonal biomass of annual forbs in creosote bush-bursage deserts Burrow construction and maintenance that creates refuges used by many commensal species (e.g., small mammals, reptiles, invertebrates), increasing local shelter availability and microhabitat heterogeneity Nutrient cycling through fecal deposition around burrow areas, concentrating nutrients and organic matter in otherwise nutrient-poor desert soils

Diet Details

Other Foods:
Spring annual forbs Perennial and subshrub forbs Grasses Succulents and cacti Desert perennial shoots and flowers

Human Interaction

Domestication Status

Wild

Agassiz's desert tortoise (Gopherus agassizii) is a wild North American species with no domestication history. People have known and used it in Indigenous cultures, collected it for pets in the 20th century, and now manage it for conservation after the 1990 listing. Human impacts include illegal collection, roads, development, grazing, invasive plants, research, rescue, relocation, and education programs.

Danger Level

Low
  • Zoonotic Salmonella exposure risk from handling reptiles or contaminated surfaces (standard reptile-associated salmonellosis risk).
  • Minor scratches or bites if handled improperly (generally uncommon due to docile behavior).
  • Indirect safety risks when people stop on roads to interact with or move tortoises (traffic hazard); only move a tortoise if legally allowed and in immediate danger.

As a Pet

Not Suitable as Pet

Legality: Usually not legal to keep Agassiz's desert tortoise (Gopherus agassizii) as a pet. Taking wild tortoises is illegal; owning or moving them is regulated. Many areas require permits or state adoption. Mojave population has federal protection.

Care Level: Expert Only

Purchase Cost: Up to $100
Lifetime Cost: $3,000 - $20,000

Economic Value

Uses:
Conservation and restoration employment Scientific research (ecology, disease, climate impacts) Education/outreach and wildlife tourism value Cultural significance (traditional knowledge)
Products:
  • No legal commercial products are associated with this protected species; economic activity is primarily indirect (conservation/research/education).

Relationships

Related Species 7

Morafka's Desert Tortoise Gopherus morafkai Shared Genus
Gopher Tortoise
Gopher Tortoise Gopherus polyphemus Shared Genus
Texas Tortoise Gopherus berlandieri Shared Genus
Bolson Tortoise Gopherus flavomarginatus Shared Genus
African Spurred Tortoise
African Spurred Tortoise Centrochelys sulcata Shared Family
Greek Tortoise Testudo graeca Shared Family
Hermann's Tortoise Testudo hermanni Shared Family

Ecological Equivalents 5

Animals that fill a similar ecological role in their ecosystem

Desert Iguana Dipsosaurus dorsalis Arid-land herbivore that relies heavily on desert annuals and perennials and uses burrows for thermoregulation. Overlaps spatially with Mojave and Sonoran desert plant communities and shares extreme heat- and drought-related physiological constraints, for example behavioral thermoregulation and seasonal activity windows.
Common Chuckwalla Sauromalus ater Large desert reptile that is primarily herbivorous (feeds on flowers and leaves), times its surface activity to avoid lethal heat, grows and reproduces slowly, and depends on shelters such as rock crevices or tortoise burrows.
Round-tailed Ground Squirrel Xerospermophilus tereticaudus Desert burrow user and consumer of seeds and green plant matter. Shares key niche elements of burrow-mediated thermal refuge and reliance on ephemeral desert productivity pulses after rainfall; burrows serve as critical microclimate buffers, similar to burrow use documented for Gopherus agassizii in Mojave systems.
American Badger Taxidea taxus Not a predator match, but a burrowing ecosystem engineer. Both species dig and modify burrows, affect soil, and provide shelter to many Mojave and Sonoran Desert animals (per USFWS).
Black-tailed Jackrabbit Lepus californicus Desert herbivore that can overlap in forage (forbs and grasses) during periods of high productivity. Ecological relationship is potential resource overlap during drought or years with low annual-plant production, and shared reliance on shrub cover and microhabitat structure for thermal regulation and predator avoidance.

“A desert tortoise can live to be over 80 years old!”

Desert tortoises are found in the United States and in Mexico. These tortoises burrow tunnels so they can go underground to cool off when the desert heat gets to be too much. The California desert tortoise eats grasses, flowers, and herbs found in their hot, dry environment. These reptiles dig grooves in the sand with their feet to catch rainwater to drink.

Infographic of Desert Tortoise

Desert Tortoise Top Facts

• Bouncing baby tortoises: The eggs laid by a tortoise are the size of ping-pong balls.
• A thirsty tortoise: After drinking rainwater, a tortoise may go as long as a year without needing more water.
• Life in a tunnel: A desert tortoise spends about 95% of its life inside tunnels beneath the sand.

Check out more incredible facts about desert tortoises.

Scientific Name

Desert Tortoise (Gopherus Agassizii)

Desert tortoise is the common name for this reptile, and Gopherus agassizii is its scientific name. This tortoise belongs to the Testudinae family, and its class is Reptilia. There is another species of desert tortoise with the scientific name Gopherus morafkai. It has a shell that’s narrower in shape than the Gopherus agassizii. Gopherus refers to the burrowing habits of this tortoise. They burrow into the ground just as actual gophers do. Agassizii is in the turtle’s name to honor Swiss zoologist Jean Louis Rodolphe Agassizii, who spent many years studying tortoises in North America.

Evolution and Origins

Desert tortoise has a long evolutionary history.

The Desert Tortoise (Gopherus agassizii) is a fascinating creature that has captured the attention of researchers and nature enthusiasts alike. Its evolutionary history is an area of great interest as scientists seek to understand how this species came into existence and how it has adapted to its harsh desert environment over time.

One thing we know for certain about the Desert Tortoise’s evolution is that it occurred over a very long period of time. Fossil evidence suggests that ancestors of today’s tortoises existed more than 50 million years ago, during the Eocene epoch. These early forms were much smaller than their modern descendants and lived in diverse environments ranging from forests to grasslands.

Over time, these ancestral tortoises evolved various adaptations that allowed them to survive in increasingly arid regions. For example, they developed thicker shells with higher domes to protect against predators and retain moisture. They also began relying on succulent plants for water and became more efficient at conserving water through urination.

The lineage leading up to Gopherus agassizii likely diverged from other tortoise species around 8-15 million years ago, during a period when geological changes created new deserts across North America. This resulted in isolated populations of tortoises adapting differently depending on their specific habitats.

Appearance and Behavior

Sonoran Desert Tortoise at Catalina State Park in Arizona.

Desert dwelling tortoises have hard shells in brown and gray.

The shell of a desert tortoise is usually brown or gray without any colorful markings on it like you would see on a Box turtle you might find in the woods. It does have a pattern of lines that separates the shell into sections or scutes. The underside of its shell is yellow or light brown.

This tortoise can be from 8 to 15 inches long and 4 to 6 inches tall. If you put a desert tortoise on a scale, it would weigh from 8 to 15 pounds. A tortoise weighing 8 pounds would weigh the same as half a bowling ball! The largest desert tortoise on record is 17 inches long and weighs 26 pounds. His name is Monster!

Desert tortoises have small black eyes and ears that cannot be seen from the outside. They have an eardrum located on their neck under a layer of scales. A desert tortoise feels the ground vibrating, and those sounds move up through its legs, shell, and into their eardrums. That’s how they hear what’s going on around them.

Whether it’s burrowing a tunnel to live in or creating a groove in the sand to catch rainwater, these tortoises do a lot of digging! They have strong front legs with sharp, sturdy nails on them that help them to make a lot of progress when breaking through the dry ground. Their scaly skin protects them from the heavy digging work they do.

A desert tortoise has a large shell with plenty of space for its lungs. Plus, its roomy shell helps this reptile to keep its body temperature normal so it can adapt to the extreme heat in the desert.

The way a desert tortoise stores water helps it to live in a hot, dry environment. After taking a big drink of rainwater, a desert tortoise can store extra water in its bladder to use whenever it needs some.

Desert tortoises like to live alone except during breeding season. However, sometimes these solitary reptiles share a tunnel with a dozen or more other tortoises, especially during the wintertime. When tortoises do form a small group, it’s called a creep. Desert tortoises are shy animals making it hard for scientists and wildlife photographers to catch a glimpse of them.

Habitat

Desert tortoises live in the southwestern part of the United States and the northwestern part of Mexico. Specifically, they live in the Mojave and Sonoran deserts. This desert environment has temperatures that sometimes go a lot higher than 105 degrees Fahrenheit, and there’s very little rainfall.

The Sonoran and the Mojave Desert tortoise survive this very hot climate by going down into tunnels during extremely hot summer weather. In fact, they go into a type of hibernation called estivation. In the summertime, desert tortoises sleep a lot to save their energy!

In the wintertime, the grasses that desert tortoises eat become very scarce. So, these reptiles move into their tunnels and go into another type of hibernation called brumation. But when springtime arrives, desert tortoises move out of their tunnels into the sunshine to eat!

Diet

What Do Tortoises Eat
Tortoises eat vegetables, fruits, plants, and fungi in the wild.

What does a desert tortoise eat? A desert tortoise eats rice grass, Bermuda grass, ryegrass, primrose, sow thistle, cactus, and wildflowers. This reptile walks slowly across the desert using its scaly, tough feet to pull dry grass out of the ground. It takes a tortoise about 20 to 30 days to digest its food!

In the wild, a desert tortoise knows what plant life to eat to survive. However, some tortoises become sick and die from eating trash left behind by humans. Balloons, plastic bags, and food containers are examples of items that are harmful to these reptiles.

Predators & Threats

Lizard Gila Monster( Heloderma suspectum) on a rock

Desert tortoises need to watch out for hungry Gila monsters.

Coyotes, skunks, ravens, foxes, and Gila monsters are all predators of the desert tortoise. These predators are likely to go after younger, more vulnerable tortoises. A desert tortoise hides in its shell or in one of its tunnels to escape a predator. Also, if it’s picked up in the mouth of a predator, it releases urine in order to make the animal let go of it. This may help the tortoise to escape the predator’s grip, but releasing urine means the tortoise has less water to drink. This can put the tortoise at risk, especially during the hot summers in the desert.

The conservation status of the desert tortoise is: Threatened. Desert tortoises are losing some of their habitats to humans who are building neighborhoods and creating more landfills in the area. Also, the tortoise is at risk when it crosses roads where vehicles travel.

Reproduction, Babies & Lifespan

Desert Tortoises lay about a dozen eggs at a time.

Reproduction

Male desert tortoises compete with each other for the attention of a female during the breeding season. One male may even push another over onto its shell to prove its strength.

The gestation period of a female desert tortoise is 3 to 4 months. She digs a nest and can lay up to 14 eggs. After laying the eggs, the female tortoise leaves them. Eggs are laid between May and July and hatch between August and October.

Babies

Once the eggs hatch, each tortoise baby, or hatchling, measures about 1.5 inches long and weighs less than a pound. The hatchlings must try to survive without their mother from birth. Many of them don’t survive because their protective shell doesn’t fully develop until they are a few years old. They must find food on their own and often fall prey to one of their many desert predators.

Lifespan

Both male and female desert tortoises can live up to 80 years old. Of course, a desert tortoise that lives in a zoo is likely to live longer than one in the wild. Living in a zoo means the tortoise doesn’t have to deal with predators and has a regular supply of food. The oldest land tortoise on record is named Jonathan. He is believed to be 185 years old!

As a desert tortoise gets older, it can suffer from various illnesses. A loss of habitat and reduced food sources can weaken a tortoise’s immune system causing it to be at risk for upper respiratory illness, shell diseases, and herpesvirus.

Population

Slowest Animals In North America

One of the reasons for tortoises’ slow speeds is because of their diet; they are herbivores, so they do not need to hunt or chase their food.

The desert tortoise population has fallen by 90% since 1980 due to habitat loss, livestock grazing, predators, and disease. Furthermore, only 1 to 5 of every 100 desert tortoise hatchlings grow to be adults. As a result, their conservation status is classified as Threatened. However, the desert tortoise was given protected status by the Endangered Species Act in 1990.

• Around 150,000 desert tortoises are living in a habitat threatened by new construction projects and trash dumping.

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How to say Desert Tortoise in ...
German
Kalifornische Gopherschildkröte
English
Desert Tortoise
French
Tortue du désert
Hungarian
Kaliforniai üregteknős
Japanese
サバクゴファーガメ
Dutch
Gopherus agassizii

Sources

  1. David Burnie, Dorling Kindersley (2011) Animal, The Definitive Visual Guide To The World's Wildlife / Accessed January 18, 2010
  2. Tom Jackson, Lorenz Books (2007) The World Encyclopedia Of Animals / Accessed January 18, 2010
  3. David Burnie, Kingfisher (2011) The Kingfisher Animal Encyclopedia / Accessed January 18, 2010
  4. Richard Mackay, University of California Press (2009) The Atlas Of Endangered Species / Accessed January 18, 2010
  5. David Burnie, Dorling Kindersley (2008) Illustrated Encyclopedia Of Animals / Accessed January 18, 2010
  6. Dorling Kindersley (2006) Dorling Kindersley Encyclopedia Of Animals / Accessed January 18, 2010
Heather Hall

About the Author

Heather Hall

Heather Hall is a writer at A-Z Animals, where her primary focus is on plants and animals. Heather has been writing and editing since 2012 and holds a Bachelor of Science in Horticulture. As a resident of the Pacific Northwest, Heather enjoys hiking, gardening, and trail running through the mountains with her dogs.

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Desert Tortoise FAQs (Frequently Asked Questions)

Coyotes, foxes, Gila monsters, badgers and road runners are all predators of the desert tortoise. However, these predators are not likely to try to eat an adult desert tortoise. An adult desert tortoise weighs a lot and is difficult to get to when it goes into its shell. Instead, these predators go after the young hatchlings because they are small and easy to capture. This is why so few hatchlings make it to adulthood. Ravens go after hatchlings and even steal the eggs from a desert tortoise’s nest. A mountain lion is one predator that will eat an adult tortoise. A mountain lion has strong jaws and is able to break through an older, more brittle shell of an adult desert tortoise.