P
Species Profile

Père David’s Deer

Elaphurus davidianus

China's rescued wetland deer

Père David’s Deer Distribution

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Endemic Species
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Found in 1 country

Size Comparison

Human 5'8"
Père David’s Deer 3 ft 8 in

Père David’s Deer stands at 65% of average human height.

Largest Deer - Pere David's Deer

At a Glance

Wild Species
Also Known As Milu, Milu deer, David's deer, Chinese milu, Elaphure
Diet Herbivore
Activity Crepuscular+
Lifespan 12 years
Weight 200 lbs
Did You Know?

All living Père David's deer descend from a small captive founder group (widely cited as 18 animals) assembled at Woburn Abbey (UK) in the 1890s.

Scientific Classification

Père David's deer (Elaphurus davidianus), also called milu, is a large cervid historically native to the wetlands and river plains of eastern China. It became extinct in the wild by the early 20th century and survived through captive herds, later enabling reintroduction programs in China.

Kingdom
Animalia
Phylum
Chordata
Class
Mammalia
Order
Artiodactyla
Family
Cervidae
Genus
Elaphurus
Species
Elaphurus davidianus

Distinguishing Features

  • Eponym honors the French missionary-naturalist Armand David (Père David).
  • Large deer with adaptations to wet habitats (often described as favoring marshy environments).
  • Unique genus (Elaphurus), distinct from Cervus (red deer) and Dama (fallow deer).

Physical Measurements

Males and females differ in size

Height
3 ft 9 in (3 ft 7 in – 3 ft 11 in)
3 ft 9 in (3 ft 7 in – 3 ft 11 in)
Length
8 ft 2 in (7 ft 10 in – 8 ft 8 in)
8 ft 4 in (7 ft 10 in – 8 ft 10 in)
Weight
441 lbs (397 lbs – 507 lbs)
287 lbs (220 lbs – 353 lbs)
Tail Length
1 ft 8 in (1 ft 6 in – 1 ft 10 in)
1 ft 8 in (1 ft 8 in – 1 ft 10 in)
Top Speed
31 mph
running

Appearance

Primary Colors
Secondary Colors
Skin Type Dense seasonal fur with pronounced winter shag; cloven, splayed hooves adapted to soft wetland ground.
Distinctive Features
  • Large wetland-associated deer with long tail ~50 cm (Nowak, 1999).
  • Head-body length 190-215 cm; shoulder height 115-122 cm (Nowak, 1999).
  • Adult mass commonly ~135-200 kg, males heavier on average (Nowak, 1999; Macdonald, 2006).
  • Broad, splayed hooves and well-developed dewclaws aid traction in marshes and mud.
  • Unique antler configuration: main beam sweeps backward with forward-pointing tines; males often show atypical antler cycle timing for deer.
  • Shaggy winter pelage and a mane/ruff on neck and shoulders are especially evident in adult males.
  • Notable conservation history: extinct in the wild by early 20th century; persisted in captive herds enabling reintroductions in China.

Sexual Dimorphism

Males are larger-bodied with heavier necks and substantially larger, more complex antlers. Females are smaller, lack antlers, and typically show a finer head-and-neck profile with less pronounced mane in winter.

  • Larger average body mass and shoulder height than females.
  • Large, multi-tined antlers with backward-sweeping main beams.
  • More pronounced neck mane/ruff, especially in winter coat.
  • Thicker neck and chest; overall more robust forequarters.
  • No antlers (antlerless).
  • Smaller, lighter-bodied build with slimmer neck.
  • Less pronounced winter mane/ruff and overall shagginess.
  • Generally subtler dorsal stripe contrast than adult males.

Did You Know?

All living Père David's deer descend from a small captive founder group (widely cited as 18 animals) assembled at Woburn Abbey (UK) in the 1890s.

It has an unusually long tail for a deer-about 50 cm-giving it a more "horse-like" silhouette than most cervids.

Adults are large: head-body length ~190-215 cm; shoulder height ~115-122 cm; males commonly ~150-200 kg and females ~100-140 kg (values vary by source and population).

A strong swimmer, it readily wades and swims in marshes and river plains, feeding on grasses and aquatic plants.

The species was extinct in the wild by the early 20th century, but reintroductions began in China in the 1980s (notably at Beijing Nanhaizi Milu Park and Jiangsu Dafeng).

Its Chinese nickname means "four not alike," reflecting a traditional belief it resembles parts of several animals rather than any single one.

Unlike most deer, its antler form is "reversed" compared with typical cervids: the main beam sweeps backward with forward-pointing tines, a shape often noted as distinctive for Elaphurus.

Unique Adaptations

  • Hoof structure suited to soft ground: splayed hooves with well-developed dewclaws increase support and traction on mud and in shallow water-useful in marsh habitats.
  • Swimming proficiency: a heavy-bodied deer that still swims well, allowing it to exploit riverine and coastal wetland mosaics that deter many terrestrial predators.
  • Distinctive antlers (genus-level trait): antler beams sweep backward with forward-pointing tines; this unusual geometry is a hallmark of the monotypic genus Elaphurus.
  • Seasonal coat shift: reddish-brown in warmer months and grayer in winter, improving camouflage across reedbeds, mudflats, and winter-dormant marsh vegetation.
  • Long tail (~50 cm): longer than in most deer, likely aiding insect defense and signaling within tall wetland vegetation.

Interesting Behaviors

  • Wetland living: spends long periods wading or standing in shallow water and muddy flats, using aquatic areas to forage and thermoregulate.
  • Herding: forms social groups; outside the rut, males and females often separate, while mixed groups become more common in the breeding season.
  • Rut displays: males engage in parallel-walking, vocalizing, scent-marking, and antler pushing/wrestling to compete for access to females.
  • Wallowing and grooming: uses mud wallows and water to cool down and reduce biting insects; vigorous body shaking after leaving water is common.
  • Foraging style: grazes grasses and sedges but also takes reeds and aquatic vegetation; often feeds with head low while moving through marsh vegetation.
  • Calf-rearing: females isolate briefly around birth, then rejoin groups; calves typically rely on hiding behavior early on (common among many cervids).

Cultural Significance

In China, Père David's deer (Elaphurus davidianus) links to imperial history because the last native herd lived in Nanyuan Imperial Hunting Park near Beijing. Nicknamed "four not alike," it is a symbol of conservation: lost from the wild, kept in zoos abroad, and returned by reintroduction programs.

Myths & Legends

"Four-Unlikes": In Chinese folk description, Père David's deer was said to resemble parts of four animals rather than any single one (wording varies by region and storyteller).

"Investiture of the Gods": In this classical Chinese mythic tradition, a strategist rides a fantastical mount called the "four-unlikes," later popularly linked in name and imagination to the deer.

Imperial park tales: Traditional anecdotes around the Nanyuan Imperial Hunting Park portray the deer as a rare royal possession-an animal seen by few commoners, lending it an aura of auspicious rarity and "forbidden" prestige.

Naming story (historical tradition): The English name commemorates Père Armand David, the French missionary-naturalist who first reported the species to Western science in the 1860s after learning of the deer kept in the imperial enclosure.

Conservation Status

EW Extinct in the Wild

Known only to survive in captivity.

Population Increasing

Protected Under

  • CITES Appendix I
  • China: National Key Protected Wild Animals (Class I)

Life Cycle

Birth 1 calf
Lifespan 12 years

Lifespan

In the Wild
8–15 years
In Captivity
12–23 years

Reproduction

Mating System Polygyny
Social Structure Harem Based
Breeding Pattern Seasonal
Fertilization Internal Fertilization
Birth Type Internal_fertilization

During the annual rut (typically June-August), males fight and guard harems, mating with multiple females. Females give birth to a single fawn after ~280-290 days' gestation and provide all parental care.

Behavior & Ecology

Social Herd Group: 12
Activity Crepuscular, Diurnal, Cathemeral
Diet Herbivore Emergent wetland grasses and sedges-especially reed- and sedge-dominated vegetation (e.g., Phragmites australis, Carex spp.)

Temperament

Generally gregarious and tolerant within herds; strong flight response when disturbed in open wetlands.
Adult males become aggressive during rut, with frequent sparring and dominance displays (Cervidae rut behavior).
Females are strongly protective around parturition; calf defense and heightened vigilance are common.
Most populations show herd living with sex segregation; activity shifts more nocturnal under human disturbance.
Longevity is highest in managed herds; captive individuals reported living >20 years (IUCN/managed-population reports).

Communication

alarm snorts
barks
grunts/moans during rut
contact calls between hind and calf
scent marking via preorbital gland secretions and urine/feces at frequently used paths
visual displays: antler posturing, parallel walking, head-lowering before sparring
tactile interactions: nudging and grooming within nursery groups
flehmen response to assess female reproductive state via urine scents

Habitat

Biomes:
Wetland Freshwater Temperate Grassland Temperate Forest
Terrain:
Plains Riverine Coastal Muddy
Elevation: Up to 656 ft 2 in

Ecological Role

Large-bodied wetland grazer/browser shaping plant community structure in floodplain marsh ecosystems.

Regulates wetland vegetation structure via grazing and selective feeding (can suppress dominance of tall emergent stands and maintain habitat heterogeneity) Nutrient cycling through dung/urine deposition, enhancing localized soil and water nutrient dynamics Facilitates seed dispersal (endozoochory of some wetland plant propagules) and seedbed creation via trampling/disturbance Creates and maintains openings/paths in dense marsh vegetation, indirectly benefiting other wetland fauna by increasing habitat patchiness

Diet Details

Other Foods:
Wetland grasses Sedges Rushes Common reed Cattails Aquatic and semi-aquatic forbs and macrophytes Shoots and leaves of riparian shrubs and trees +1

Human Interaction

Domestication Status

Wild

Père David's deer (Elaphurus davidianus) is wild, not domesticated. After extinction in the wild, it survived only in captive care. Animals were kept in the Imperial Hunting Park (Nanyuan) near Beijing until about 1900. Individuals sent to Europe, including Woburn Abbey (UK), formed the founder stock for breeding and allowed reintroduction to China from the 1980s.

Danger Level

Moderate
  • Injury from kicking/striking when restrained or cornered (powerful hind legs typical of large cervids)
  • Goring or lacerations from antlers (especially adult males during rut)
  • Crush injuries from body mass (adult males commonly ~150-200 kg) during handling/transport
  • Zoonotic/biological risks shared with other deer when in close contact (e.g., parasites; bacterial pathogens), mitigated by veterinary screening and hygiene
  • Vehicle-collision risk where free-ranging or reintroduced populations occur

As a Pet

Not Suitable as Pet

Legality: Père David's deer (Elaphurus davidianus) are generally not legal or practical as private pets. Many places treat them as large exotic wildlife, needing permits, special licenses, secure facilities, vet care, and transport approvals; laws vary.

Care Level: Expert Only

Purchase Cost: $2,000 - $10,000
Lifetime Cost: $50,000 - $150,000

Economic Value

Uses:
Conservation breeding and reintroduction (high value; core use) Zoo exhibition, education, and research (high) Ecotourism in reserves/rewilded landscapes (moderate) Limited/private game-ranch style ownership where legal (low; rare) Cultural/heritage value in China (moderate)
Products:
  • Non-consumptive: zoo display/education programming, conservation genetics value, ecotourism revenues
  • Research outputs: reproductive biology, population management, wetland restoration indicator species
  • (Not typical/usually discouraged for this species) consumptive products common in other cervids-venison, hides, antlers/velvet-are generally not primary or appropriate economic products for milu conservation populations

Relationships

Related Species 8

Elaphurus bifurcatus Elaphurus bifurcatus Shared Genus
Eld's deer Rucervus eldii Shared Family
Swamp deer Rucervus duvaucelii Shared Family
Red deer
Red deer Cervus elaphus Shared Family
Sika deer
Sika deer Cervus nippon Shared Family
Fallow deer
Fallow deer Dama dama Shared Family
Sambar
Sambar Rusa unicolor Shared Family
Roe deer
Roe deer Capreolus capreolus Shared Family

Ecological Equivalents 5

Animals that fill a similar ecological role in their ecosystem

Swamp deer Rucervus duvaucelii Occurs in floodplain and riverine grasslands and marsh edges, grazing tall grasses and sedges and using seasonal floods. Père David's deer (Elaphurus davidianus) is similarly adapted to soft, wet ground and is large (about 1.15–1.22 m at the shoulder, 100–200 kg).
Marsh deer Blastocerus dichotomus Convergent wetland specialization in marshes, flooded savannas, and riparian wetlands, with similar locomotor and foraging demands—moving through shallow water and grazing on emergent macrophytes. Ecological analog despite occurring on different continents (South America vs. East Asia).
Sika deer
Sika deer Cervus nippon Père David's deer and Sika deer overlap in East Asian lowland habitats such as river valleys and reedbeds. Both eat grasses and forbs, use edge habitats, form groups, feed in open areas, and rest in cover.
Chinese water deer
Chinese water deer Hydropotes inermis Shares use of lowland riverine and reedbed habitats in eastern China. Although smaller and possessing tusks rather than antlers (unlike Pere David's deer), it occupies similar riparian wetland and edge habitats and is subject to similar disturbance and predation pressures in fragmented lowlands.
Moose
Moose Alces alces Comparable browser-grazer that uses aquatic and emergent vegetation along pond and lake margins and in riverine wetlands and frequently wades. Ecological similarity is based on wetland foraging strategy rather than close taxonomy or geography.
Père David’s deer is an extraordinary creature. Sporting massive antlers on a horse-shaped head, it possesses a donkey-like tail and cow-like hooves, complete with webbing between its toes for swimming.

“A Pere David’s deer has webbing between its toes and is an excellent swimmer”

Pere David’s deer live in large groups. They are herbivores that mostly eat grass. The average lifespan of this deer is 18 years. Most female Pere David’s deer have just one baby or fawn. A fawn stands up and nurses its mother within hours of being born.

5 Pere David’s Deer Facts

Largest Deer - Pere David's Deer

Père David’s deer originate from the northeastern and east-central regions of China.

  • Pere David’s deer are from the northeastern and east-central parts of China
  • They live in swamps and marshlands
  • These deer have spread hooves with webbing to help them swim
  • They are social animals living together in large groups.
  • These deer have a reddish coat in the summer and a gray coat in the wintertime

Scientific Name

The Père David's deer (Elaphurus davidianus), also known as the milu or elaphure, Adult male

Though commonly known as Pere David’s deer, this animal is formally referred to by its scientific name, Elaphurus davidianus.

While Pere David’s deer is this animal’s common name, its scientific name is Elaphurus davidianus. The Latin term Elaphurus means it belongs to the Cervidae (deer) family.

Davidianus refers to the French zoologist and Catholic Priest Father Armand David who discovered this deer in China. The family of this deer is Cervidae and its class is Mammalia. In France, the word for Father is Pere so the literal translation of this mammal’s name is Father David’s deer.

The Chinese have another name for the Pere David’s deer. The word is sibuxiang and means ‘four not alike’. This refers to the fact that this mammal has hooves like a cow, the neck of a camel, the tail of a donkey, and the antlers of a deer. In other words, this deer has the features of four animals all rolled into one!

Evolution and Origins

The Père David’s deer (Elaphurus davidianus), alternatively called milu or elaphure, belongs to the deer species found in China’s subtropical river valleys. Its diet primarily consists of grass and aquatic vegetation. It stands as the sole surviving species within the Elaphurus genus.

Additionally, the Père David’s deer faced a big threat because people hunted them a lot and their homes disappeared. Europeans took the remaining deer to zoos in Europe before 1912. Almost 100 years later, in 1985, China got 38 deer back to reintroducing them into the wild.

Further, the Père David’s deer stands out among deer. It has large antlers on a horse-shaped head, a donkey-like tail, and cow-like hooves, but with webbed toes for swimming.

Appearance and Behavior

A stunning Milu Deer also lnown as Pere David's Deer) (Elaphurus davidianus) grazing in a pasture.

The coat of a Pere David’s deer is reddish-brown and features a black stripe across its shoulder.

In the summertime, a Pere David’s deer has a reddish-brown coat with a black stripe on its shoulder. But, in the winter, its coat turns grayish brown. These colors help to camouflage the deer during different seasons. Male Pere David’s deer have antlers that measure 21 to 31 inches in length.

Antlers that are 31 inches long are equal in length to 2 stacked bowling pins. Male deer use their antlers to fight other males when competing for females during mating season. These deer may rear up on their hind legs and clash their antlers together in combat.

A Pere David’s deer is 6 ½ to a little over 7 feet long. A 7-foot-long deer is as long as a large Christmas tree. Also, this deer can weigh from 298 lbs to 441 lbs A 441 lbs Pere David’s deer is equal in weight to half of a full-grown horse. While 441 lbs is the heaviest weight of a Pere David’s deer, a Whitetail deer can grow to be a little over 500 lbs.

The Pere David’s deer have hooves, but they are different from the hooves of other types of deer. Most deer have hooves that are narrow with the toes pressed close together. Alternatively, a Pere David’s deer has hooves that are spread out with webbing between their toes. Why? These hooves help to move Pere David’s deer through the water as they swim in swamps and marshes. Many of these deer spend a lot of time standing in water that can go as high as their shoulders.

The unique design of this deer’s hooves is great for swimming, but they don’t particularly add to its speed. A Pere David’s deer can only run about 18 miles per hour. Compare this to a Whitetail deer that can reach speeds of up to 30 miles per hour or a Reindeer that can run 50 miles per hour!

These deer are social, outgoing mammals and like to live in large groups sometimes called a herd, gang, or mob. When Pere David’s deer were plentiful, there may have been dozens or even hundreds of deer in a herd. Plus, living in a herd provides protection from a predator such as a tiger. Once the deer start running, a tiger has a challenging time finding and separating one deer to attack. In addition, a group of running deer can injure a tiger that’s trying to enter the herd.

Habitat

Pere David’s deer are from northeastern and east central China. They are originally from a subtropical climate living in marshlands and around swamps. Their webbed hooves allow them to spend a lot of time swimming. Their hooves also offer them support while walking in muddy, soggy areas.

Diet

What do Pere David’s deer eat? They are herbivores and eat a diet of mostly grass. However, if grass is in short supply, especially during the wintertime, they will eat aquatic plants that grow in and around the marshlands.

Pere David’s deer instinctively know what types of grasses and plants to eat. But, if they eat grass or other vegetation that’s been contaminated with chemicals or pesticides they can become very ill or die.

Predators and Threats

couple male elaphure deer burling in water

The Pere David’s deer holds the official conservation status of being Extinct in the Wild.

The official conservation status of the Pere David’s deer is Extinct in the Wild. Back in the late 1800s, there were just a few of these deer left in China. Their population had been greatly reduced by people hunting and eating them.

A small group of these deer belonged to the Emperor of China named Tonghzi. But, a flood knocked down a fence where the deer were kept, and they escaped. Those deer were hunted and eaten by peasants and soldiers in the area. So, when they lived in the wild, their main predator was man. Tigers also preyed on Pere David’s deer.

Today, there’s a relatively small number of Pere David’s deer living in captivity in zoos and sanctuaries. Efforts are being made to increase the population of these deer while they are living in protected areas.

Reproduction, Babies, and Lifespan

Pere David's deer (Elaphurus davidianus), also known as the milu or elaphure

Pere David’s deer have a mating season in June. Males mate with one female or more.

Mating season is in June for this deer. A male Pere David’s deer joins a group of females to mate with one or more. During this time, the male deer is likely to fight with other male deer to protect the group of females he is with. Male deer hit or box each other with their antlers as a way of fighting. The stronger male wins.

A female’s gestation period is about 9 months and she gives live birth to a baby, also called a fawn in April or May. Most female Pere David’s deer have one fawn. Having two fawns at a time is rare. Fawns weigh between 25 lbs and 29 lbs at birth and start growing quickly!

Like other fawns, these deer are born with white spots on their coat. The spots disappear as they grow older. A fawn can see when it’s born but can’t walk right away. However, a fawn struggles to get up and put weight on its legs almost immediately. This makes a lot of sense. If the newborn fawn stayed on the ground in the wild, it would be vulnerable to predators such as tigers or corsac foxes.

A fawn nurses its mother until they are weaned and begin to eat grass with the older deer. A fawn stays with its mother for about 14 months when it’s ready to survive without help from her.

Pere David’s deer live to be about 18 years old in the wild. The oldest a Pere David’s deer has lived is 23 years and that was in captivity where these deer are well-cared for.

As these deer grow older they’re prone to diseases affecting their muscle tissue also known as myopathy. Incidentally, this type of myopathy is similar to equine myopathy occurring in horses.

Population

Though Pere David’s deer’s official conservation status is Extinct in the wild, there are some exceptions. The efforts made by conservationists to increase the population have begun to work. Some of these deer have been released back into the wild in the hopes they will breed and increase the population even more. Also, there are still many Pere David’s deer being cared for in zoos and other sanctuaries. The population is estimated at around 2000.

One of the reasons for the slow population growth of this mammal is a female deer only has one baby per litter. But, if the population of Pere David’s deer continues to grow, it may receive a new, updated conservation status.

FAQ

Is the Pere David’s deer extinct?

Officially, Pere David’s deer are listed as Extinct in the Wild. But, this doesn’t mean there are none in existence anywhere. Breeding programs have allowed the release of some of these deer back into the wild. In addition, there are Pere David’s deer on display at various zoos in China, the United States, and elsewhere.

How many Pere David’s deer are left?

Approximately 2,000 of these deer are left in the world. Conservationists are working to increase this deer population each year.

How did the Pere David’s deer become extinct in China?

The Pere David’s deer became extinct in the wild in China because they were regularly hunted for food or sport. Also, the loss of their wetland habitat contributed to the massive drop in this deer population. Road construction is one thing that is taking wetland space away from Pere David’s deer and other animals that live there.

What does a Pere David’s deer eat?

This deer eats grass on a daily basis. However, if grass is not plentiful at various times of the year, they will eat aquatic plants growing in the marshlands.

Who discovered Pere David’s deer?

This deer gets its name from the Catholic priest and zoologist/botanist who discovered it. His name was Father Armand David. He discovered this deer as well as many other species of animals and plants on his expeditions to China in the mid-1800s. He was sent there to document these species so others could learn more about them.

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Rebecca Bales

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Père David’s Deer FAQs (Frequently Asked Questions)

Pere Davids Deer are Herbivores, meaning they eat plants.