A
Species Profile

Anchovy

Engraulidae

Small Fish, Big Impact
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School of Swimming Anchovies

At a Glance

Family Overview This page covers the Anchovy family as a group. Stats below are general traits shared across the family.
Also Known As Anchoa, Anchois, Acciughe, Boquerones, Anchoveta, Dilis, Katakuchi-iwashi
Diet Filter Feeder
Activity Diurnal+
Lifespan 3 years
Weight 0.3 lbs
Status Not Evaluated
Did You Know?

Family-wide size spans ~2.5 cm to ~40 cm, from tiny coastal species to larger, herringlike anchovies.

Scientific Classification

Family Overview "Anchovy" is not a single species but represents an entire family containing multiple species.

Anchovies are small, schooling, plankton-feeding marine fishes that serve as key forage for larger fish, seabirds, and marine mammals; many species support large commercial fisheries.

Kingdom
Animalia
Phylum
Chordata
Class
Actinopterygii
Order
Clupeiformes
Family
Engraulidae

Distinguishing Features

  • Small, slender silvery body adapted for schooling
  • Single dorsal fin; generally soft-rayed clupeiform body plan
  • Distinctly large mouth in many species, with the upper jaw often extending behind the eye
  • Filter-feeding or particulate-feeding on plankton; important mid-trophic ‘forage fish’ role

Physical Measurements

Males and females differ in size

Length
♂ 5 in (2 in – 1 ft 2 in)
♀ 5 in (1 in – 1 ft 1 in)
Weight
♂ 0 lbs (0 lbs – 0 lbs)
♀ 0 lbs (0 lbs – 1 lbs)
Top Speed
19 mph
Short bursts 10–30 km/h

Appearance

Primary Colors
Secondary Colors
Skin Type Thin, delicate skin with small cycloid scales (often easily shed/deciduous) and a noticeable mucus layer; overall body is streamlined and laterally compressed for sustained schooling swimming.
Distinctive Features
  • Family-level size range (generalization across Engraulidae): ~4-40 cm total length from the smallest micro-anchovies to the largest long-bodied anchovies; most species are in the ~8-20 cm range.
  • Lifespan range (varies by species and region): commonly ~1-4 years in small, fast-growing coastal pelagic species, with some species/populations reaching ~6-8+ years under favorable conditions; strong year-class variability is typical.
  • Elongate, laterally compressed body with a single dorsal fin set mid-body and a forked to moderately forked tail; overall 'forage-fish' shape optimized for schooling.
  • Relatively large mouth for body size (upper jaw often extending behind the eye in many taxa), reflecting planktivory and, in some species, the ability to take larger prey items.
  • Silvery flanks and countershading aid in predator avoidance in open/coastal pelagic waters; reflectance and darkness of the dorsum often shift with water clarity, depth use, and condition.
  • Ecology/behavior generalization: strongly schooling and highly abundant where conditions are favorable; forms dense surface or midwater aggregations that are key prey for larger fishes, seabirds, and marine mammals (major forage-fish role).
  • Feeding generalization: primarily plankton-feeding (copepods and other zooplankton; phytoplankton/microplankton in some contexts), using ram-feeding and/or filter-feeding tendencies; diet and feeding mode vary with species, size, and local plankton composition.
  • Habitat generalization: mostly coastal pelagic, commonly on continental shelves and nearshore waters; many species regularly use bays/estuaries and tolerate broad salinity ranges, while others are more offshore; a minority occur in brackish-to-freshwater systems.
  • Reproduction generalization: typically high fecundity with pelagic eggs/larvae and seasonal or multi-batch spawning; timing and duration vary widely with latitude, upwelling intensity, and local oceanographic conditions.
  • Fishery relevance: many Engraulidae support large commercial reduction and/or food fisheries; populations can fluctuate strongly with ocean climate (e.g., upwelling strength, temperature anomalies), producing boom-bust dynamics and variable body condition.

Did You Know?

Family-wide size spans ~2.5 cm to ~40 cm, from tiny coastal species to larger, herringlike anchovies.

Many species form immense schools-dense enough to be seen by satellites via seabird and predator activity in some upwelling regions.

They are classic forage fish: a key link moving plankton energy into larger fish, seabirds, and marine mammals.

Several species tolerate brackish water and routinely enter estuaries and lagoons, while others stay fully marine.

Reproduction is often "batch spawning": multiple spawn events across a season, helping populations rebound quickly when conditions are good.

Anchovy fisheries can swing dramatically with ocean climate (e.g., warming events and changes in upwelling), reshaping ecosystems and economies.

Not all "anchovies" in markets are the same species-many regional fisheries harvest different Engraulidae with similar roles and culinary uses.

Unique Adaptations

  • Protrusible, forward-projecting upper jaw and wide gape that help capture tiny prey efficiently in the water column.
  • Fine, often numerous gill rakers that enable filtering small zooplankton; raker spacing and reliance on filtering vary across species.
  • Silvery, reflective flanks for countershading and "school shimmer" camouflage in open water.
  • High fecundity and relatively fast growth (family-wide), supporting boom-bust dynamics tied to ocean productivity.
  • Tolerance (in several lineages) to changing salinity and turbidity, enabling use of estuaries and coastal lagoons as feeding or nursery areas.

Interesting Behaviors

  • Mass schooling and synchronized turning to reduce individual predation risk; school size and tightness vary by species, habitat, and predator pressure.
  • Coastal pelagic lifestyle: many species track plankton-rich fronts, river plumes, or upwelling zones; some stay nearshore, others range farther offshore seasonally.
  • Plankton feeding by filtering or picking: some species rely more on filter-feeding with gill rakers, others more on selective biting-often shifting with prey size and density.
  • Diel (day-night) changes in depth and schooling: schools may tighten or change depth with light levels and predator activity; patterns differ among regions and species.
  • Spawning strategies vary widely: some spawn in open coastal waters, others near estuaries; egg and larval drift can strongly influence where juveniles end up.
  • Predator-driven behavior: rapid, wave-like "flash expansions" and sudden dives are common when attacked by tuna, seabirds, or marine mammals.

Cultural Significance

Engraulidae (anchovies) support important small-pelagic fisheries and coastal foods. Salted anchovies are a staple in the Mediterranean; Southeast Asian fish sauces use them; South American upwelling fuels big catches for food and fishmeal. They shaped old trade and port economies.

Myths & Legends

Anchovies (Engraulidae): the name 'anchovy' came into English from Spanish and Portuguese words and may link to a Basque word meaning anchovy, showing the fish's long role in Bay of Biscay fishing.

Roman culinary tradition: ancient writers describe fermented fish sauces made from small pelagic fish-an enduring cultural ancestor of later Mediterranean anchovy condiments and sauces.

Medieval and early modern European coastal practice: salted "poor man's fish" (including anchovy-type preserves) became tied to Lenten/fasting food traditions, embedding anchovies in seasonal religious food calendars.

Peru's rise of industrial anchovy fishing in the 20th century became a national economic story, a warning about El Niño years, collapse risk, and how ocean climate can shape society.

Mediterranean port-city stories link anchovies (Engraulidae) to sailors and long voyages. Dishes like Provence onion-and-anchovy tart and Italian anchovy recipes are shown as born from preserving fish at sea.

Conservation Status

NE Not Evaluated (higher-taxon/family level; IUCN assessments are generally made at the species level)

Has not yet been evaluated against the criteria.

Population Unknown

You might be looking for:

European anchovy

22%

Engraulis encrasicolus

Common in the eastern Atlantic and Mediterranean; widely used in cuisine.

Peruvian anchoveta

20%

Engraulis ringens

Southeast Pacific; among the world’s most heavily harvested forage fishes.

Northern anchovy

16%

Engraulis mordax

Eastern Pacific (California Current); important forage species.

Japanese anchovy

14%

Engraulis japonicus

Northwest Pacific; major fishery species in East Asia.

Indian anchovy

10%

Stolephorus indicus

Indo-West Pacific; common coastal anchovy in regional fisheries.

Life Cycle

Birth 30000 frys
Lifespan 3 years

Lifespan

In the Wild
1–10 years
In Captivity
1–36 years

Reproduction

Mating System Polygynandry
Social Structure Aggregation Group
Breeding Pattern Transient
Fertilization Broadcast Spawning
Birth Type Broadcast_spawning

Anchovies (Engraulidae) are small, schooling fish that reproduce by mass, broadcast spawning (polygynandry). Many males and females release eggs and sperm in groups; no pair bonds or parental care; brief spawning events.

Behavior & Ecology

Social School Group: 10000
Activity Diurnal, Crepuscular, Cathemeral
Diet Filter Feeder Copepods (dominant, energy-rich zooplankton prey for many Engraulidae, though composition varies by region and season)
Seasonal Migratory 186 mi

Temperament

Strongly social and aggregation-prone; individuals rely on group cohesion and rapid alignment to reduce predation risk (a key ecological role as forage fish).
Generally non-territorial; competition is mostly exploitative (tracking plankton patches) rather than aggressive defense.
Startle-prone and risk-averse: abrupt, synchronized escape responses are common; school density often increases under threat.
Feeding/space-use flexibility varies across the family: many are coastal marine and surface-oriented, while others regularly use estuaries and some occupy brackish-to-freshwater reaches; schooling remains common across these habitats.
Ecology generalization with variation: most are plankton-feeding (filtering/particulate feeding on phyto- and zooplankton), but diet breadth shifts with size, local productivity, and season; some species take larger zooplankton or fish larvae more frequently.
Measurement range across the family (smallest to largest members): roughly ~2-40 cm total length (most species are small-bodied, commonly <20 cm).
Lifespan range across species: typically short-lived, roughly ~1-8 years (many mature early and experience high natural and fishing mortality; longer lifespans occur in some larger/less-exploited populations).

Communication

No specialized vocal communication is well-established across Engraulidae; any sounds are generally incidental E.g., from rapid swimming/feeding or schooling turbulence
Vision-based alignment and neighbor-position tracking within schools Daylight and clear-water conditions strengthen cohesion
Lateral line detection of hydrodynamic cues to maintain spacing, match speed/direction, and coordinate collective turns-especially important in low light or turbid water.
Chemical cues (kairomones/alarm substances) likely contribute to predator awareness and group responses, though strength and documentation vary by species.
Collective displays and movement patterns (e.g., tightening, flash expansion, coordinated diving/rising) function as group-level signaling/anti-predator tactics; expression varies with predator type and habitat.

Habitat

Coastal Open Ocean Seabed/Benthic Estuary Mangrove Wetland River/Stream Lake Rocky Shore Kelp Forest Coral Reef +5
Terrain:
Coastal Island Sandy Muddy
Elevation: Up to 1312 ft 4 in

Ecological Role

Forage-fish plankton consumer linking lower trophic levels (phyto/zooplankton) to higher predators; a key mid-trophic component in many coastal upwelling and shelf ecosystems, with strong spatial/seasonal variability across species and regions.

Transfers primary production to higher trophic levels (supports predatory fishes, seabirds, and marine mammals) Regulates plankton communities via intense grazing during blooms Nutrient cycling and redistribution through schooling, excretion, and vertical movements Supports major commercial and artisanal fisheries; provides prey base that stabilizes/structures food webs

Diet Details

Main Prey:
Zooplankton Krill Cladocerans Ostracods Larval crustaceans Fish eggs and larvae Polychaete larvae +1
Other Foods:
Phytoplankton Microalgae and suspended plant-like particles

Human Interaction

Domestication Status

Wild

Anchovies (Engraulidae) are wild-caught fish used by people through capture fisheries, not domestication. Humans fish them along coasts and in industrial fleets, cure and salt them, and use them as bait or fishmeal. No true domesticated lineage exists; some are kept for study, but regular breeding is rare. Impacts range from small nets to large reduction fisheries.

Danger Level

Low
  • Food-related risks: allergic reactions in sensitive individuals; histamine (scombroid-like) poisoning can occur if fish are mishandled/temperature-abused after capture; foodborne illness risk if undercooked or improperly preserved.
  • Physical hazards: minor punctures or abrasions during handling of nets/gear; choking hazard from small bones if eaten whole (varies with preparation).
  • Occupational/indirect risks: injuries associated with fishing operations (slips, entanglement, cuts) rather than from the fish themselves.

As a Pet

Not Suitable as Pet

Legality: Anchovies (Engraulidae) are usually legal to keep where marine fish are allowed, but local fishing rules (size limits, seasons, baitfish rules), marine protected area limits, import and animal welfare laws may apply.

Care Level: Expert Only

Purchase Cost: Up to $20
Lifetime Cost: $500 - $5,000

Economic Value

Uses:
Commercial capture fisheries (food) Reduction fisheries (fishmeal/fish oil) Bait fisheries Processing/food manufacturing Ecosystem support for higher-value fisheries (forage base)
Products:
  • fresh/chilled anchovies (region-dependent)
  • canned anchovies
  • salted/cured anchovies
  • anchovy paste
  • fish sauce/fermented anchovy products (in regions where used)
  • bait (hook-and-line, traps, recreational fishing)
  • fishmeal and fish oil (including omega-3 sources)
  • animal feed ingredients and fertilizers (via fishmeal byproducts)

Relationships

Predators 9

Tunas Thunnus spp.
Mackerel Scomber spp.
Bonitos and skipjacks Sarda spp.; Katsuwonus pelamis
Hake Merluccius
Cod
Cod Gadidae
Seabirds Laridae; Sternidae; Pelecanidae; Phalacrocoracidae
Dolphin
Dolphin Delphinidae
Seals and sea lions Pinnipedia
Sharks
Sharks Selachimorpha

Related Species 12

European anchovy Engraulis encrasicolus Shared Family
Peruvian anchoveta Engraulis ringens Shared Family
Northern anchovy Engraulis mordax Shared Family
Japanese anchovy Engraulis japonicus Shared Family
Indian anchovy Stolephorus indicus Shared Family
Pacific anchoveta Cetengraulis mysticetus Shared Family
Atlantic anchoveta Cetengraulis edentulus Shared Family
Bay anchovy Anchoa mitchilli Shared Family
Striped anchovy Anchoa hepsetus Shared Family
Dussumier's hairfin anchovy Coilia dussumieri Shared Family
Grenadier anchovy Coilia ramcarati Shared Family
Gangetic hairfin anchovy Setipinna phasa Shared Family

Ecological Equivalents 5

Animals that fill a similar ecological role in their ecosystem

Sardines
Sardines Sardinops spp. Small, schooling, pelagic forage fish that filter-feed on plankton and support major reduction fisheries. They often co-occur and fill similar mid-trophic roles.
Herring
Herring Clupea spp. Schooling clupeiform fishes with comparable plankton-feeding ecology and predator-prey importance in coastal and shelf ecosystems.
Menhaden Brevoortia spp. Highly abundant, plankton-feeding forage fish that form dense schools; share ecosystem function and fishery uses, including as bait and in reduction fisheries.
Capelin Mallotus villosus Not closely related, but ecologically similar as a small, schooling, lipid-rich forage fish that links plankton to larger fish, seabirds, and marine mammals.
Sand lance Ammodytes spp. Similar role as schooling forage fish heavily targeted by predators, but many sand lances burrow in the substrate and are less strictly plankton-filtering.

Types of Anchovy

24

Explore 24 recognized types of anchovy

European anchovy Engraulis encrasicolus
Peruvian anchoveta Engraulis ringens
Northern anchovy Engraulis mordax
Japanese anchovy Engraulis japonicus
Cape anchovy Engraulis capensis
Australian anchovy Engraulis australis
Indian anchovy Stolephorus indicus
Commerson's anchovy Stolephorus commersonnii
Buccaneer anchovy Stolephorus buccaneeri
Spined anchovy Stolephorus tri
Shorthead anchovy Encrasicholina heteroloba
Dotted anchovy Encrasicholina punctifer
Bay anchovy Anchoa mitchilli
Striped anchovy Anchoa hepsetus
Bigeye anchovy Anchoa lamprotaenia
Pacific anchoveta Cetengraulis mysticetus
Atlantic anchoveta Cetengraulis edentulus
Dussumier's hairfin anchovy Coilia dussumieri
Grenadier anchovy Coilia ramcarati
Japanese grenadier anchovy Coilia nasus
Phasa / Gangetic hairfin anchovy Setipinna phasa
Scaly hairfin anchovy Setipinna taty
Moustached thryssa Thryssa mystax
Hamilton's thryssa Thryssa hamiltonii

Quick Take

  • Anchovies have maintained a global presence for 15 million years since the Miocene epoch.
  • The upper jaw extension behind the eyes complicates species identification during classification.
  • Surprisingly, select anchovy species have successfully transitioned into the fresh waters of South America.
  • Using crab paste is necessary to lure anchovies to the surface during the deepwater trawling stage.

Anchovies are small green forage fish. These fish are very common and high in nutrition and belong to the family Engraulidae. There are more than 100 species of anchovies — most of which are found in marine waters. However, some of these small fish are also found in brackish waters, while others can also exist in the fresh waters of South America.

Anchovies are fish that are mostly distinguished by their large mouths. Their upper jaw extends behind their eyes. Besides that, a long, pig-like snout extends beyond the tips of their lower jaws.

A comprehensive infographic about anchovies featuring anatomical diagrams, a global habitat map, evolutionary timelines, and culinary uses.
More than just a controversial pizza topping, these 15-million-year-old survivors are the hidden engine of the world's oceans. See how they've outlasted epochs to become the ultimate marine powerhouse. © A-Z Animals

Incredible Facts

  • These fish are easier to catch on a full moon night since their bodies glisten under the moonlight. They are canned and salted right after, and sometimes dried for eating purposes.
  • Crab paste is known to be the best bait for catching these fish.
  • November 12th is celebrated as National Pizza with the Works Except Anchovies Day.
  • The freshest anchovies have black irises instead of reddish ones. They are also often confused with Sardines that have white flesh.

Classification and Scientific Name

Anchovies are fish that belong to the kingdom Animalia and phylum Chordata. Their class is Actinopterygii, and they come from the order of Clupeiformes. These small fish belong to the family Engraulidae. Engraulidae comes from the Greek word Engraulis, which means anchovy. However, the name possibly comes from the word γρυλίζω, meaning “to grumble.”

Species

According to sources, there are 144 different species worldwide. These different species are distributed into 17 genera. The majority of these species differentiate as a result of their location, although there are a few ways in which they distinguish themselves.

Some of the common species include:

  • European anchovy (Engraulis encrasicolus): These anchovies are one of the most common species to consume, and they are predominantly located in the Mediterranean Sea and the North Sea. They often have a silver underbelly, spawning through the spring and summer months.
  • Peruvian anchoveta (Engraulis ringens): This species is often found in the southeast Pacific Ocean, and it is harvested by the ton.
  • Indian anchovy (Stolephorus indicus): The Indian anchovy also goes by the name Hardenberg’s anchovy. It can be used as either live bait or as dead bait.
  • Northern anchovy (Engraulis mordax): Not to be mistaken for the European anchovy, this species is typically found in two separate parts of the United States, spanning across Oregon, Washington, and California.

Evolution

School of anchovies

Anchovies have been on Earth for millions of years.

Anchovies have been around for millions of years, and their evolution is believed to be closely linked to the evolution of other species in the same family, such as herrings and sardines. It is thought that anchovies evolved from a common ancestor that lived during the Miocene epoch, some 15 million years ago. Since then, they have adapted to inhabit many different environments across the world’s oceans and seas.

Anchovies are now considered one of the most abundant fish species. Anchovy populations tend to fluctuate due to changes in ocean temperatures caused by climate change, making them vulnerable if not managed properly. Despite this vulnerability, anchovies remain one of our planet’s most important food sources due to their high nutritional value, which includes essential omega-3 fatty acids and protein content — meaning they are here to stay for generations more yet!

Anchovies vs. Sardines

Anchovies, as a result of curing, often have dark, reddish-gray flesh, while Sardines have white flesh. Sardines have a slightly protruding lower jaw, while the upper jaw of Anchovies extends behind their eyes.

Sardines are generally larger than anchovies, though sizes can vary depending on the species

The main way that these two types of fish differ is in their taste. Anchovies tend to make any dish saltier. However, for individuals who want a meatier taste that is a little more mellow and subdued, then sardines may be the right choice.

Sardines belong to the Clupeidae family, while anchovies belong to the Engraulidae family.

Appearance

School of anchovies in the Atlantic Ocean

Anchovies are small green fish with longitudinal stripes of blue and silver.

These are small green fish. They are slender and streamlined and resemble herrings a lot. Usually, they are categorized by large mouths.

Their upper jaws extend behind their eyes, and a pointed pig-like snout extends beyond the lower jaw. They have a straight lateral line.

These fish have large scales that can be easily detached. An adult fish is about 5 to 6 inches in size. Even the heaviest of these fish does not weigh more than a few ounces.

Distribution, Population, and Habitat

These fish are found in several oceans around the world. They live in scattered spots throughout these oceans. However, they are mostly concentrated in temperate waters.

They are very rare or are completely absent in areas with extreme temperatures. That is why it is difficult to find them in very cold or very warm waters. The European species is especially abundant in the Black Sea, Alboran Sea, and Aegean Sea.

These fish are usually found in saltwater. However, some of them can also exist in brackish waters. They are pelagic, meaning that they thrive best in open oceans. In brackish waters, they exist in estuaries and bays.

According to a study, as of 2015, the estimated population is 20,700 metric tons.

Predators and Prey

Sailfish

Sailfish rely on anchovies and sardines for food.

These fish, especially the bay anchovies, have bluefish, striped bass, and weakfish as predators. Apart from that, seabirds also feed on them, including the bay anchovy. Other predators of these fish include sharks, salmon, halibut, and other fish. They, in turn, eat recently hatched fish as well as plankton.

What Eats Anchovies?

Anchovies are an important food source for many animals, including birds such as seagulls, penguins, and pelicans. Many marine mammals, such as dolphins, seals, and whales, rely on anchovies for food. Large predatory fish like tuna and mackerel, and other smaller fish like sardines and herring, all eat anchovies. Several types of reptiles, like sea turtles and crustaceans such as crabs and lobsters, also eat anchovies. Anchovies provide essential nutrients to all these creatures since they contain high levels of calcium, phosphorus, magnesium, potassium, sodium chloride (salt), iodine, trace elements/minerals needed for metabolic activities. Thus, anchovies play a key role in maintaining the balance of life in oceans around the world.

What Do Anchovies Eat?

These fish feed mainly on plankton as well as recently hatched fish.

Reproduction and Lifespan

Peruvian anchoveta

Anchovies reproduce by spawning.

Much like herring, the fish reproduce by spawning. Males and females gather in large groups and release their eggs and sperm into the water.

The sperm are known to fertilize the eggs outside the bodies of the fish. The eggs, once fertilized, float in water columns until they hatch.

The gestation period and the time that it usually takes for the babies to reach sexual maturity vary from species to species. They rarely live past four years of age.

Fishing and Cooking

Most of these fish are caught by Deepwater trawlers. Once caught, most of these fish are canned and salted. After, they are turned into a paste which is later used to make sauces. Crab paste is a common bait used to bring anchovies to the surface.

Further, they are considered to be a magic ingredient in dishes. They have a savory and briny flavor. Usually, they are stir-fried in tomato sauce or shallow-fried in olive oil.

People also love these small fish as toppings on their pizza or butter them up in their toast as an easy snack idea. They are sometimes dried for eating purposes. Dried fish also makes great snacks.

Population and Conservation

School of anchovies

Overfishing is an issue for the declining anchovy populations.

The most recent population study of anchovies puts their population around 20,000 metric tons.

Anchovies are classified as a ‘Least Concern’ species on the IUCN Red List, meaning they have healthy populations and their conservation status is considered low risk. However, there are still several threats to anchovy populations around the world. Overfishing of anchovies has been an issue for centuries, with some stocks becoming depleted due to unsustainable fishing practices. Anchovies can also be threatened by water pollution from agricultural runoff or oil spills, which contaminate their habitats and food sources. Additionally, climate change is causing ocean temperatures to rise in many areas, which affects fish reproduction cycles and migratory patterns. It is important that we continue to monitor anchovy populations so we can ensure sustainable fisheries management and protect these important fish from further harm or a decline in numbers.

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Sources

  1. New World Encyclopedia / Accessed December 24, 2020
  2. Wikipedia / Accessed December 24, 2020
  3. Mobile cuisine / Accessed December 24, 2020
  4. Food and Wine / Accessed December 24, 2020
  5. Nature Gate / Accessed December 24, 2020
  6. Animals / Accessed December 24, 2020
  7. The Spruce Eats / Accessed December 24, 2020
Heather Hall

About the Author

Heather Hall

Heather Hall is a writer at A-Z Animals, where her primary focus is on plants and animals. Heather has been writing and editing since 2012 and holds a Bachelor of Science in Horticulture. As a resident of the Pacific Northwest, Heather enjoys hiking, gardening, and trail running through the mountains with her dogs.

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Anchovy FAQs (Frequently Asked Questions)

Anchovies are found in different oceans around the world. They live in scattered spots throughout these oceans. However, they are mostly concentrated in temperate waters.