S
Species Profile

Sand Dollar

Clypeasteroida

Coins of the seafloor, built to burrow
k_lmnop/Shutterstock.com

Sand Dollar Distribution

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This map shows coastal regions where Sand Dollar are found.

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Size Comparison

Human 5'8"
Sand Dollar 1 in

Sand Dollar stands at 1% of average human height.

What Do Sand Dollars Eat - Sand Dollar In Hand

At a Glance

Order Overview This page covers the Sand Dollar order as a group. Stats below are general traits shared across the order.
Also Known As Sea biscuit, Sea cookie, Cake urchin, Keyhole urchin, Mermaid's money, Mermaid's coin
Activity Cathemeral+
Lifespan 8 years
Weight 1.5 lbs
Status Not Evaluated
Did You Know?

"Sand dollar" is a common-name group; the order Clypeasteroida also includes thicker "sea biscuits/cake urchins."

Scientific Classification

Order Overview "Sand Dollar" is not a single species but represents an entire order containing multiple species.

Sand dollars are flattened echinoids (sea urchin relatives) adapted for life on or within sandy seafloors. They have a rigid internal skeleton (test), short movable spines, tube feet for feeding/respiration, and a characteristic petal-like pattern on the upper surface.

Kingdom
Animalia
Phylum
Echinodermata
Class
Echinoidea
Order
Clypeasteroida

Distinguishing Features

  • Flattened, disk- to oval-shaped test (skeleton) with five-petaled pattern on the upper surface (petaloids)
  • Very short spines giving a velvety texture when alive
  • Mouth on the underside; anus typically near the edge or on the underside depending on species
  • Often burrow in sand; some species have characteristic holes/slots (e.g., Mellita)

Physical Measurements

Males and females differ in size

Height
1 in (0 in – 3 in)
1 in (0 in – 2 in)
Length
4 in (1 in – 12 in)
5 in (1 in – 10 in)
Weight
0 lbs (0 lbs – 4 lbs)
0 lbs (0 lbs – 2 lbs)
Top Speed
0 mph
crawling

Appearance

Primary Colors
Secondary Colors
Skin Type Rigid calcareous test covered by very short, fine, movable spines (velvety); tube feet emerge through pores for feeding/respiration; living surface often traps sand and detritus.
Distinctive Features
  • Order-level form spans very flattened, disk-like sand dollars to thicker, biscuit-like/cake forms within Clypeasteroida.
  • Adult size range across the order: ~2-3 cm to ~20-25+ cm test diameter, with thickness from a few millimeters to several centimeters.
  • Lifespan across species commonly several years; roughly ~3 to 20+ years depending on size, habitat, and predation.
  • Typically inhabit sandy or mixed-sediment seafloors from surf/shallow subtidal to deeper shelf settings; depth range varies widely by species and region.
  • Many species burrow or partially bury, orienting to waves/currents; others sit on the surface, especially thicker forms.
  • Feeding is commonly deposit- or suspension-feeding using spines, cilia, and tube feet; diet and particle size preferences vary among lineages.
  • Often show abrasion-resistant, sand-shedding spines; empty tests are frequently white/cream from bleaching.
  • Five-fold symmetry is expressed as a flower-like petaloid pattern on the upper surface; visibility varies with spine cover and sediment.

Did You Know?

"Sand dollar" is a common-name group; the order Clypeasteroida also includes thicker "sea biscuits/cake urchins."

Across the order, adult size ranges from ~1-2 cm (tiny dwarf sand dollars) to ~25-30+ cm across (largest sea biscuits).

Many species show a 5-petaled pattern on top; it reflects specialized tube feet used for gas exchange.

Some sand dollars have slots (lunules) through the body that help reduce lift and improve stability in surf zones.

They're echinoderms: their bodies are built on a 5-part plan even when they look "coin-shaped."

A living sand dollar's surface is covered in short, velvety spines that move sand and food particles like a conveyor belt.

Beach "sand dollars" are often just the bleached internal skeleton (test); living ones are usually darker and bristly.

Unique Adaptations

  • Flattened (or thickened) rigid test: thin sand dollars reduce drag and allow burial; thicker sea biscuits are more resistant to crushing and often live on/just under sand.
  • Petaloid "flower" fields: paired rows of pores house tube feet specialized for respiration; the pattern is a hallmark across much of the order.
  • Short, highly coordinated spines: act as legs, shovels, and particle conveyors, moving both the animal and a continuous layer of sand over the body.
  • Lunules (in many sand dollars): slots/perforations through the test that reduce hydrodynamic lift and help keep the animal anchored in strong surge.
  • Fine particle-handling: ciliated spines and mucus help trap tiny food particles; feeding mode shifts with current speed and sediment type.
  • Reinforced internal architecture: the test's internal struts and plates provide strength while staying relatively lightweight for sandy habitats.

Interesting Behaviors

  • Burrowing and self-burying: many species rapidly shimmy into sand using coordinated spine strokes; depth and speed vary by species and grain size.
  • Feeding flexibility: commonly deposit-feeding on organic films and microalgae, suspension-feeding on fine particles in moving water, or doing both depending on currents and habitat.
  • Current-orientation: in wave-swept habitats some sand dollars align or partially stand with flow to optimize feeding and avoid being flipped; others stay deeply buried.
  • Group living: some species form dense beds (sometimes thousands per area) that can stabilize sediment and influence local food availability; others are more solitary.
  • Broadcast spawning is common (eggs/sperm released into water), producing planktonic larvae; timing and frequency vary with temperature and season, and a few lineages show more localized reproductive strategies.
  • Predator responses: burial, spine motion, and "hunkering down" are typical; predators include sea stars, fishes, crabs, and large gastropods-risk varies strongly by region and depth.

Cultural Significance

Sand dollar (Clypeasteroida) are famous beach finds used in crafts and souvenirs. Their petal pattern and inner pieces are often seen as Christian symbols (doves, a star, Easter designs) and used to teach children. Large groups can show healthy sandy sea bottoms in coastal education and citizen science.

Myths & Legends

In a modern coastal Christian story, the Sand Dollar's petal-like pattern is said to hide Nativity and Crucifixion symbols. When the shell breaks, it reveals five small 'doves' (ossicles) showing peace and resurrection.

Mermaid's coin / sea money (Atlantic and Caribbean beach folklore): sand dollars are described as coins dropped by mermaids or sea spirits, washed ashore as tokens of luck or safe travel.

Poseidon/Neptune's currency (classical-inspired seaside tale): beachcombers in some coastal traditions call them the sea god's coins, linking their coin shape to the ocean's ruler and good fortune on voyages.

In many beach cultures, finding an intact sand dollar Clypeasteroida is seen as a sign of luck, fortune, or a wish granted; people often give the sand dollar to someone else to share luck.

Conservation Status

NE Not Evaluated (order-level hub; IUCN assesses most taxa at species level)

Has not yet been evaluated against the criteria.

Population Unknown

You might be looking for:

Pacific sand dollar

24%

Dendraster excentricus

Common sand dollar of the west coast of North America; often found in surf-zone sandy bottoms.

Common sand dollar

22%

Echinarachnius parma

North Atlantic species; frequently washed ashore and widely referenced in guides and collections.

Keyhole sand dollar (five-hole sand dollar)

18%

Mellita quinquiesperforata

Western Atlantic/Caribbean sand dollar with distinct perforations (“holes”).

Cake urchins (often called sand dollars)

14%

Clypeaster

Thicker, more domed relatives within Clypeasteroida; commonly grouped under “sand dollar” in casual usage.

Life Cycle

Birth 500000 larvas
Lifespan 8 years

Lifespan

In the Wild
3–20 years
In Captivity
1–10 years

Reproduction

Mating System Promiscuity
Social Structure Aggregation Group
Breeding Pattern Not Applicable
Fertilization Broadcast Spawning
Birth Type Broadcast_spawning

Across sand dollars, sexes are separate and adults usually live solitary or in loose benthic aggregations. Many species synchronize seasonal spawning, releasing eggs and sperm into the water; fertilization is external and mates are multiple, with no pair bonds or parental care.

Behavior & Ecology

Social Aggregation Group: 50
Activity Cathemeral, Nocturnal, Diurnal
Diet Detritivore Diatom-rich organic detritus and microbial biofilm associated with fine sand

Temperament

Non-aggressive; lacks territorial defense and relies on burrowing, camouflage, and spines.
Generally sedentary to slow-moving; activity peaks vary with currents, sediment, and predation risk.
Often tolerant of close neighbors in dense beds; spacing can still occur via sediment disturbance.
Predator avoidance ranges from shallow burial to deep burial depending on species and substrate.
Across the order, social interactions are mostly incidental; true cooperative behavior is absent.

Communication

none detected
Chemical cues in seawater E.g., spawning synchrony, conspecific presence
Tactile contact via spines and tube feet during crowding or jostling in shifting sand
Hydrodynamic cues and vibration/sediment disturbance sensing from nearby movement
Gamete release timing influenced by environmental signals (temperature, tides, currents), varying by species

Habitat

Seabed/Benthic Coastal Beach Open Ocean Coral Reef Kelp Forest Estuary +1
Biomes:
Terrain:
Coastal Island Sandy Muddy
Elevation: Up to 6561 ft 8 in

Ecological Role

Benthic deposit-feeding echinoids that process sandy sediments and recycle organic matter in nearshore soft-bottom ecosystems

bioturbation and sediment mixing (reworking surface layers) nutrient remineralization and enhancement of benthic microbial productivity sediment cleaning/sorting that can influence grain-size distribution and organic content at micro-scales transfer of detrital energy to higher trophic levels as prey for fishes, crabs, and other predators contribution to benthic-pelagic coupling by intercepting resuspended organic particles in dynamic environments

Diet Details

Other Foods:
Organic detritus Particulate organic matter Microalgae and diatom films Bacterial biofilms Suspended organic particles trapped from the water column

Human Interaction

Domestication Status

Wild

Clypeasteroida (sand dollars and sea biscuits) are wild marine animals with no domesticated lineages. People mostly find them by beachcombing and collect dead tests. Scientists and educators collect some for study. A few enter the marine aquarium trade but usually do not live long in captivity.

Danger Level

Low
  • Minor skin puncture/irritation from spines (generally short and not highly injurious compared with many sea urchins)
  • Cuts from sharp test fragments when handling dried/broken specimens
  • Ethical/legal risk: taking live animals can violate local wildlife or protected-area regulations

As a Pet

Not Suitable as Pet

Legality: Laws differ by country, state, and protected area. Collecting live sand dollars/sea biscuits is banned or limited in many coastal places and marine protected areas; permits may be needed. Dead, beach-worn tests are often allowed, but check local rules.

Care Level: Expert Only

Purchase Cost: Up to $100
Lifetime Cost: $300 - $5,000

Economic Value

Uses:
Curio/souvenir trade (mostly dead tests) Education and research specimens Public aquaria/exhibits (limited, specialized) Ecosystem services (bioturbation and sediment mixing)
Products:
  • Dried sand dollar/sea biscuit tests sold as souvenirs and craft materials
  • Teaching specimens for anatomy, development, and marine ecology
  • Research use in studies of echinoderm development, biomechanics of tests/spines, and benthic ecology

Relationships

Related Species 6

Keyhole sand dollars Mellitidae Shared Family
Pacific sand dollars Dendrasteridae Shared Family
Northern sand dollars Echinarachniidae Shared Family
Sea biscuits / large sand dollars Clypeasteridae Shared Family
Indo-Pacific sand dollars Arachnoididae Shared Family
Sand dollars and sea biscuits Laganidae Shared Family

Ecological Equivalents 4

Animals that fill a similar ecological role in their ecosystem

Heart urchins Spatangoida Occupy similar sandy, soft-bottom habitats and often burrow. Both are deposit feeders that use spines and tube feet to move sediment and funnel organic particles, though heart urchins are typically more deeply infaunal and less disk-flattened.
Sea cucumbers Holothuroidea Share the role of sediment processors on sandy bottoms, ingesting or collecting organic-rich particles. They differ in body plan and feeding structures but overlap strongly in detritus-based energy pathways.
Burrowing bivalves Bivalvia Co-occur on sandy flats and in shallow subtidal zones and rely on organic matter from the sediment and water column. Sand dollars are mainly surface/subsurface deposit feeders, while many bivalves are suspension feeders; both occupy a comparable soft-bottom infaunal/epifaunal niche.
Sand stars Astropecten spp. Live and hunt on sandy substrates and spend time partially buried. Relationship reflects ecological similarity (sand-habitat specialists) and predator-prey overlap, as some sea stars prey on small or buried echinoids.

Types of Sand Dollar

10

Explore 10 recognized types of sand dollar

Pacific sand dollar Dendraster excentricus
Northern sand dollar Echinarachnius parma
Five-holed (keyhole) sand dollar Mellita quinquiesperforata
Rosy sea biscuit Clypeaster rosaceus
Cake urchin / sea biscuit Clypeaster subdepressus
Six-holed sand dollar Leodia sexiesperforata
Michelin's sand dollar Encope michelini
Notched sand dollar Encope emarginata
Common Indo-Pacific sand dollar Arachnoides placenta
Sea biscuit (Indo-Pacific) Clypeaster humilis

The skeletons of these animals are popular with collectors. When they are alive, Sand Dollars actually have gray or purple spines and fine hairs covering their bodies. They move along the sea floor and bury themselves in the sand. When they wash onto the shore, they are usually dead and make great additions to a seashell collection.

Sand Dollar Facts

  • The larval form of Sand Dollars can clone themselves and continue to grow and develop as two distinct animals.
  • There are 250 or more species within the Sand Dollar order, Clypeasteroida.
  • Most species of Sand Dollars move along the ocean floor using their spines and cilia.
  • A few species of fish, including Cod and Flounder, eat Sand Dollars. These fish can even get through the hard skeleton of the Sand Dollar.
  • While Sand Dollars are not endangered, collecting live specimens is illegal in most areas. This is to help preserve and protect their natural habitat and the ecosystem there.
sand dollar, white

When found on the beach, Sand Dollars are dead. They are bleached white, and the cilia are no longer visible.

Classification and Scientific Name

The well-known sand dollar is actually an entire scientific order, known as Clypeasteroida. There are at least 250 species that belong to multiple suborders and families. Many share similarities, but color, size, and location can all vary.

They are part of the superorder Gnathostomata. This includes other sea urchins that are all irregularly shaped and have a central mouth on their underside. The class Echinoidea includes all sea urchins, over 900 species total. Most have round shells with hard spines all over them. Most sea urchins eat algae and are very useful in aquariums to keep the algae growth in check.

Sand dollars are part of the phylum Echinodermata. Members of this phylum are known as Echinoderms. This translates to “echino-,” which means hedgehog, and “-derm,” which means skin. While not all members of the phylum have spikes or spines like sea urchins, they all have five-pointed body structures and usually boast some sort of spiny surface. There are around 7,000 species within the phylum, making it the largest marine phylum in the animal kingdom.

Sand Dollar Appearance

Sand Dollars are usually between three and four inches across. They are shaped as irregular circles, measured by their diameter. Their skeleton is called a test and is arranged in five points. This is the portion that most collectors see on the beach. While Sand Dollars are alive, their test is covered with soft spines and cilia, fine hairs. This helps the Sand Dollar move under the water.

When you find a Sand Dollar on the beach, chances are its spines and cilia will no longer be present. Instead, the test is bleached to a white color. It still has a five-point structure design. This design is actually made of five symmetrical pairs of lines. They are the pores in the skeleton that allow the Sand Dollar to exchange gases. When they are alive in the ocean, Sand Dollars are purplish-gray, with the color varying depending on the species. This color comes from the spines and cilia on the surface of their skeletons.

The Sand Dollar’s mouth is located on the bottom at the center. The Sand Dollar’s anus is located to the side, which is unique among sea urchins. Some Sand Dollar species also have slits in their skeleton, called lunules. These remain even after the Sand Dollar has died and its skeleton washes up on the beach. Lunules help them embed more deeply in the sand without getting washed away by the current or ocean waves.

sand dollar

The cilia that help the Sand Dollar move are visible in this photo.

Distribution, Population, and Habitat

Sand Dollars are common in warm waters as well as temperate areas. Many species live off the coasts of Central America and South America. Some also live as far north as Florida and the eastern coast of the United States. Sand Dollars are also found on the Pacific coast, from Alaska all the way to Mexico.

They live in shallow water as well as deeper ocean habitats over 100 feet. As long as they have food to eat and sand to burrow in, Sand Dollars are happy. They bury themselves in the sand and often live in groups, both for reproduction and because they congregate where food is plentiful.

Sand Dollars are not endangered, but people are still discouraged from collecting or trawling for them. In fact, in most areas, taking a live Sand Dollar specimen is illegal. Trawling on the sea floor can also be illegal. This environment is very fragile and supports a variety of ocean life. The Sand Dollars are just one of many types of sea animals that live on the ocean floor. While they may not be in danger of declining population numbers, changes in their habitat can have disastrous consequences for other, more vulnerable species.

Predators and Prey

Sand Dollar spines are used for movement rather than defense or offense. Their hard exoskeleton provides some deterrent for potential predators. However, fish such as Cod and Flounder do eat Sand Dollars. Humans can also become predators for Sand Dollars, taking specimens from the ocean for collections. This is illegal in many areas, however.

Sand Dollars feast on algae and other small particles in the ocean water. They can use their spines to move along the sea floor to get to additional patches of algae. Some species turn on their side to get algae that are floating in the water. This is less common, however. They also sometimes eat crustacean larvae.

Reproduction and Lifespan

Sand Dollars can be male or female. They reproduce externally. This means that both male and female Sand Dollars release gametes into the water, where fertilization occurs. The young are called larvae, which then grow and develop into adult Sand Dollars.

Sand Dollar larvae can also clone themselves when they feel threatened. They divide into two distinct specimens, and each continues to grow and develop. These clones end up being smaller than the original larvae. Scientists think that this may help them escape the notice of predators better due to their smaller size.

Sand Dollars in Fishing and Cooking

It is illegal in most areas to fish for or collect live Sand Dollars. This is due to the fragile nature of their environment more than the risk of extinction. Sand Dollars are favorites among collectors because of their flower-like design on the front and stark white skeleton. You can usually take Sand Dollar skeletons that you find on the beach home with you, although you should always double-check the local regulations.

Some legends say that Sand Dollars are actually the currency used by mermaids or in the mythical city of Atlantis. It has also been associated with the Star of Bethlehem or an Easter Lily in Christianity.

Bakers can replicate the pattern of Sand Dollars by making cookies that also feature a five-pointed design on the front. Sugar cookies with almonds arranged in the same pattern are a fun way to give a nod to Sand Dollars at your next get-together.

sand dollars

Sand dollars are very popular among seashell collectors.

Similar Sea Urchins to Sand Dollars

Sea Stars: These also have a five-point structure, although they do not possess the same kind of exoskeleton. These relatives of the Sand Dollar also make popular ocean specimens in your collection, although make sure that you only collect dead sea stars.

Sea Cucumber: These animals also live on the sandy sea floor and have spiny, leathery skin. They are shaped like a cucumber that can move, which gives them their name.

View all 391 animals that start with S

Sources

  1. New York Times / Accessed September 27, 2022
  2. Monterey Bay Aquarium / Accessed September 27, 2022
  3. Oregon Coast Aquarium / Accessed September 27, 2022
Katie Melynn Wood

About the Author

Katie Melynn Wood

Katie is a freelance writer and teaching artist specializing in home, lifestyle, and family topics. Her work has appeared in At Ease Magazine, PEOPLE, and The Spruce, among others. When she is not writing, Katie teaches creative writing with the Apex Arts Magnet Program in Anne Arundel County, Maryland. You can follow Katie @katiemelynnwriter.
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Sand Dollar FAQs (Frequently Asked Questions)

Sand Dollars can only survive for a few minutes out of the water. If you see a Sand Dollar washed up on the beach and it is stark white without any spines, it is already dead and you are actually looking at its skeleton. If it still has spines attached, get it back into the water right away.