C
Species Profile

Caribou

Rangifer tarandus

Antlers, Arctic, and endless miles.
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Caribou Distribution

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Invasive Species
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Size Comparison

Human 5'8"
Caribou 3 ft 11 in

Caribou stands at 69% of average human height.

Caribou Migration

At a Glance

Wild Species
Also Known As Tuktu
Diet Herbivore
Activity Cathemeral+
Lifespan 10 years
Weight 300 lbs
Status Vulnerable
Did You Know?

Both sexes usually grow antlers; males typically shed soon after the autumn rut, while many females keep antlers through winter (helps them access food at snowy feeding sites).

Scientific Classification

A circumpolar deer species adapted to cold climates; North American populations are commonly called caribou. Notable for large seasonal migrations in some herds and for both sexes typically growing antlers.

Kingdom
Animalia
Phylum
Chordata
Class
Mammalia
Order
Artiodactyla
Family
Cervidae
Genus
Rangifer
Species
Rangifer tarandus

Distinguishing Features

  • Both males and females usually bear antlers (females typically smaller)
  • Broad, splayed hooves adapted for snow, ice, and marshy ground
  • Dense insulating coat; often pale/gray-brown with lighter neck and rump
  • Nasal adaptations that warm incoming cold air
  • Strong migratory behavior in many tundra populations

Physical Measurements

Males and females differ in size

Height
3 ft 11 in (2 ft 9 in – 4 ft 11 in)
3 ft 5 in (2 ft 7 in – 3 ft 11 in)
Length
6 ft 3 in (5 ft 4 in – 7 ft)
6 ft 2 in (5 ft 2 in – 7 ft 5 in)
Weight
397 lbs (220 lbs – 701 lbs)
198 lbs (110 lbs – 331 lbs)
Tail Length
5 in (2 in – 8 in)
6 in (3 in – 8 in)
Top Speed
50 mph
Short sprint about 80 km/h

Appearance

Primary Colors
Secondary Colors
Skin Type Thick mammalian skin covered by a dense, double-layer coat: insulating underfur plus hollow guard hairs that increase buoyancy and reduce heat loss; pronounced seasonal molt (winter coat longer and more insulating).
Distinctive Features
  • Both male and female caribou usually grow antlers with many tines. Antlers vary by sex, age, and population. Males tend to have larger, heavier antlers with a shovel-like brow tine.
  • Large, crescent-shaped hooves of Caribou (Rangifer tarandus) have sharp edges for cutting ice. Their footpads change with seasons—harder and exposed in winter for traction, softer and wider in summer for boggy ground.
  • Specialized nasal passages with complex turbinates that warm and humidify cold air and recover heat/moisture during exhalation-supporting cold-climate endurance (reported in physiological studies of Rangifer and summarized in mammalogy references).
  • Dense neck mane/ruff (especially in winter) and a relatively short tail; the bright rump patch is conspicuous when running.
  • Migration and herd behavior often change caribou appearance: tundra migrants show big seasonal fat and coat changes, while some boreal/woodland groups stay put and change less.
  • Adults vary by subspecies and region: body length about 1.6–2.1 m, shoulder height 0.9–1.4 m; females usually 80–120 kg, males about 110–210+ kg.
  • Longevity: commonly ~10-15 years in the wild; up to ~20 years reported in captivity under managed conditions (typical cervid pattern; reported in species summaries such as ADW and mammalogy references).
  • Caribou antler cycle links to breeding and winter feeding competition: males usually shed after the autumn rut; females often keep antlers through winter and shed after calving, affecting herd look and sex ID.
  • Note: In North America they are called caribou; in Eurasia called reindeer. Both are the same species, Rangifer tarandus. Local ecotypes, subspecies, and domestication change coat color and size.
  • Conservation status differs by population: some boreal and woodland caribou have steep declines, while many tundra herds stay larger. This changes herd sex and age makeup and how herds look.

Sexual Dimorphism

Caribou (Rangifer tarandus) show sexual dimorphism mainly in body size, antler size/shape, and when antlers are shed. Both sexes usually have antlers; males are larger with heavier, complex antlers, females are smaller with slimmer antlers and keep them later into winter.

  • Larger body size on average: commonly heavier and taller than females within the same population; adult males often ~110-210+ kg depending on ecotype (broad compiled ranges in mammalogy references and species accounts).
  • Antlers usually larger/heavier with more pronounced tines; antler growth supports dominance during the rut.
  • Typically shed antlers soon after the rut (late autumn to early winter in many populations), so adult males may be antlerless for much of winter.
  • Smaller body size: often ~80-120 kg depending on ecotype; generally more lightly built than males.
  • Antlers present in most adult females but smaller and more slender; useful for competition at winter feeding sites.
  • Often retain antlers through winter and shed after calving in spring/early summer, so antlered individuals in late winter groups are frequently females and juveniles (context-dependent).

Did You Know?

Both sexes usually grow antlers; males typically shed soon after the autumn rut, while many females keep antlers through winter (helps them access food at snowy feeding sites).

Gestation is about 228-234 days (~7.5 months); calves are commonly ~5-8 kg at birth and can stand and nurse within hours.

Seasonal migrations can span hundreds to >1,000 km annually in some tundra herds (e.g., Alaska-Yukon's Porcupine herd is among the best-known long-distance migrants).

Reindeer/caribou can see into the ultraviolet spectrum; studies show their eyes shift seasonally to improve contrast in snowy Arctic light (e.g., Stokkan et al., 2013, Proc. R. Soc. B).

Their hooves change with the seasons: in winter the hoof pads shrink and harden for traction on ice and crusted snow; in summer pads soften for grip on wet tundra.

They are exceptional swimmers, routinely crossing wide, cold rivers and lakes during migrations.

Milk is extremely energy-rich compared with many other deer-reindeer milk is reported around ~18% fat in husbandry references-supporting rapid calf growth in short Arctic summers.

Unique Adaptations

  • Cold-specialized coat: dense underfur plus hollow guard hairs trap air for insulation and buoyancy; the coat also sheds seasonally to match temperature and insect pressure.
  • Seasonally tuned hooves: winter-hardening hoof pads improve traction on ice and help break crusted snow; summer-soft pads increase grip on wet, uneven tundra.
  • Nasal heat and moisture recovery: enlarged nasal turbinates and complex passages warm inhaled air and recapture heat/water from exhaled air-key for breathing in extreme cold without excessive water loss.
  • Wide, splayed hooves act like snowshoes, spreading weight on soft snow and tundra; they also function as efficient "shovels" when cratering.
  • Low-light/UV vision: ocular adaptations (including seasonal changes in the reflective layer of the eye) enhance contrast in Arctic winter light and make UV-dark objects stand out against snow (reported in peer-reviewed work such as Stokkan et al., 2013).
  • Energy strategy for short summers: rapid summer foraging and fat storage support survival and reproduction through long winters when diets can rely heavily on lichens and sparse vegetation.

Interesting Behaviors

  • Mass migration & route fidelity: some populations follow traditional calving and wintering routes year after year; other ecotypes (e.g., many boreal forest woodland caribou) are more sedentary and widely dispersed.
  • Calving strategy: many tundra herds synchronize births within a short window in late spring/early summer, often using open tundra or windswept areas that can reduce predator encounter rates.
  • "Cratering" for food: individuals dig through snow with their forehooves to reach lichens and buried vegetation; groups may reuse and expand feeding craters.
  • Rut behavior (typically Sep-Oct): males compete via displays, herding, vocalizations (including grunts/clicks), and antler sparring; dominant males guard groups of females.
  • Insect-avoidance aggregations: during peak mosquitoes/warble flies, animals bunch tightly and move to windy ridges, snowfields, or coastal areas to reduce biting pressure.
  • River/lake crossings: herds may stage, then pour into water in large numbers; crossings can be timed with migration pulses and calving movements.
  • Herd structure varies by region: large mixed-sex herds are common in migratory tundra systems, while many boreal woodland caribou occur in small groups or as dispersed individuals, especially outside the rut.

Cultural Significance

Reindeer (caribou, Rangifer tarandus) are central to Arctic peoples for food, clothing, shelter, tools, and trade. Sami and Nenets reindeer herding shapes land use and traditional knowledge. They are listed as Least Concern, but some regional herds (e.g., Canadian boreal woodland) are threatened by habitat loss, changing predators, and industry.

Myths & Legends

Caribou (Reindeer) (Rangifer tarandus) are central to who the Sami are and to their ways of living; seasonal herding moves shape yearly work and community life, and reindeer appear in many Sami stories and beliefs.

In Inuit and other Arctic stories, versions of 'The Woman Who Married a Caribou' explain human-caribou family ties, hunting taboos, and how to treat caribou (Rangifer tarandus) with respect so herds return.

Nenets reindeer-herding lore includes tales in which reindeer are sacred companions given by higher powers; proper ritual respect toward the herd is portrayed as essential for safe travel and survival on the tundra.

Athabaskan/Dene and nearby northern traditions tell caribou change stories where animals and humans spoke the same language; the tales teach giving back: take only what you need and honor herds to renew.

Modern global folklore links reindeer to winter gift-bringers: the flying-reindeer motif popularized in 19th-century literature (notably "A Visit from St. Nicholas," 1823) has become a widely shared seasonal legend rooted in northern reindeer imagery.

Conservation Status

VU Vulnerable

Facing a high risk of extinction in the wild.

Population Decreasing

Protected Under

  • IUCN Red List: Rangifer tarandus assessed as Vulnerable (VU) with a decreasing population trend (latest published global assessment for the species).
  • Harvest is regulated across much of the range via national/provincial/territorial wildlife laws (e.g., permit/quota/season frameworks in Canada, Alaska/USA, Norway, Sweden, Finland, and Russia), but protection strength varies widely by herd/population.
  • Many key habitats occur within protected areas (national parks, reserves) in multiple range states; however, protected-area coverage is often incomplete relative to migration routes and seasonal ranges.
  • Notably at-risk units within the broader caribou/reindeer complex are frequently listed under subnational instruments (e.g., in Canada, several designatable units/ecotypes such as Boreal Caribou and Peary Caribou are listed under the federal Species at Risk Act), reflecting that conservation risk varies greatly among populations even while the species is assessed globally as VU.
  • CITES: Rangifer tarandus is not generally listed in the CITES Appendices (international trade controls are therefore not the primary conservation tool at the species level).

Life Cycle

Birth 1 calf
Lifespan 10 years

Lifespan

In the Wild
4–18 years
In Captivity
10–23 years

Reproduction

Mating System Polygyny
Social Structure Harem Based
Breeding Pattern Seasonal
Fertilization Internal Fertilization
Birth Type Internal_fertilization

Rangifer tarandus (caribou, reindeer) breeds in a short autumn rut (Sept-Oct). Males fight, form and guard harems (polygyny), and dominant bulls do most mating. Gestation ~228-234 days; single calves arrive May-June. Females breed ~16-28 months and rear calves alone.

Behavior & Ecology

Social Herd Group: 30
Activity Cathemeral, Crepuscular
Diet Herbivore Reindeer lichens (Cladonia spp.; especially the Cladonia rangiferina / C. arbuscula group)
Seasonal Migratory 839 mi

Temperament

Highly social and generally non-territorial; group cohesion is flexible (fission-fusion) with frequent joining/splitting tied to forage, insects, and movement demands (Leader-Williams 1988).
Typically vigilant and flight-prone when disturbed; reliance on early detection and rapid movement is common in open habitats.
Dominance interactions occur, especially during rut and at concentrated resources; males show increased aggression and competition during breeding season (Leader-Williams 1988).
Seasonally variable spacing: tighter aggregation is common during migration and in response to insects/predators, while more dispersed foraging groups are common when conditions allow (e.g., Bergerud 1974; Fancy & White 1987).

Communication

Low grunts and nasal snorts used in close-range social contexts E.g., during herding, agitation, or rut
Calf bleats/contact calls and maternal response calls supporting cow-calf cohesion.
Rut-associated male vocalizations (short grunts/calls) during tending and contests.
Acoustic non-vocal signaling: prominent clicking sounds produced by tendons/sesamoid structures in the feet during walking, aiding contact/coordination in low visibility and potentially signaling movement rate within groups Commonly documented for Rangifer
Olfactory communication: urine/feces and glandular scents used for individual/sexual status assessment, especially during rut E.g., investigation of scent marks and anogenital sniffing; Leader-Williams 1988
Visual displays: antler and head-posture displays, threat gestures, and chasing/parallel walking during dominance interactions Notably in rut and at feeding sites
Tactile contact: cow-calf nuzzling and close following; occasional body contact during crowding/migration.

Habitat

Biomes:
Tundra Boreal Forest (Taiga) Alpine Wetland Temperate Rainforest Temperate Forest
Terrain:
Plains Plateau Hilly Mountainous Valley Coastal Island Riverine Rocky +3
Elevation: Up to 8202 ft 1 in

Ecological Role

Large-bodied circumpolar tundra-boreal herbivore (often migratory) that strongly influences plant community composition and functions as a key prey base for northern predators.

Regulates tundra and boreal vegetation via grazing/browsing (including strong effects on lichen mats and shrub/graminoid balance) Nutrient cycling and fertilization through feces and urine, concentrating nutrients along migration routes and seasonal ranges Physical ecosystem engineering via trampling and snow 'cratering,' which can expose vegetation and alter soil/snow microhabitats Seed and propagule dispersal (external attachment and endozoochory for some plant material) Supports food webs by sustaining predators (e.g., wolves) and providing carrion for scavengers, especially during harsh winters

Diet Details

Other Foods:
Reindeer lichens Lichens Graminoids Forbs Willow Birch browse Dwarf shrubs Mosses and other bryophytes Mushrooms Aquatic and emergent tundra vegetation +4

Human Interaction

Domestication Status

Semi domesticated

Rangifer tarandus (caribou/reindeer) were domesticated in northern Eurasia only in the last ~2,000–3,000 years, with large-scale herding rising in the past ~500–1,000 years among Sami, Nenets and others. Many animals remain partly wild and breed with wild herds. Humans use them for meat, hides, transport, milk, and face hunting, conservation, conflict, and health problems.

Danger Level

Moderate
  • Physical injury from kicks or antler goring-risk increases with adult males during rut and females defending calves
  • Handling/transport injuries (large-bodied cervid; can panic, charge, or strike fencing/people)
  • Roadway hazards: collisions with vehicles in regions where herds cross roads
  • Zoonotic/occupational exposure risks for hunters/herders (e.g., Brucella spp. in some Rangifer systems; parasitic infections), plus general carcass-handling risks
  • Public-health/regulatory risk interface with cervid diseases (e.g., chronic wasting disease concerns affecting movement/keeping of captive cervids in some areas)

As a Pet

Not Suitable as Pet

Legality: Not usually legal or practical as a household pet. Rangifer tarandus is often kept as livestock or captive deer. In the U.S. and Canada laws vary; permits, ID, disease testing (CWD), transport limits, fencing, zoning, or bans may apply.

Care Level: Expert Only

Purchase Cost: $1,500 - $8,000
Lifetime Cost: $25,000 - $90,000

Economic Value

Uses:
Subsistence food and cultural use (wild caribou) Pastoral livestock herding (semi-domesticated reindeer) Meat production Hides/leather and fur Antlers (craft/ornament) and velvet antler products Draft/transport (sled pulling) and pack use Milk (regional/limited) Ecotourism and cultural tourism Seasonal exhibition/entertainment (regulated displays)
Products:
  • meat (fresh, dried, smoked)
  • fat and marrow (food use)
  • hides for clothing, footwear, bedding, tents
  • leather goods
  • antler handicrafts/tools
  • velvet antler (supplement markets in some regions)
  • sled/transport services (working animals)
  • limited dairy products (where practiced)

Relationships

Ecological Equivalents 6

Animals that fill a similar ecological role in their ecosystem

Muskox
Muskox Ovibos moschatus Co-occurs with many tundra caribou herds in Arctic and alpine ecosystems. Both are cold-adapted, wide-ranging northern ungulates that rely heavily on graminoids and sedges and forage through snow in winter, and both are important prey for wolves and bears in tundra food webs.
Moose
Moose Alces alces Shares boreal (taiga) landscapes with many woodland caribou; both are large northern cervids influenced by snow depth, seasonal shifts in forage availability, and predation risk (notably wolves). Moose are more browse-specialized, while caribou often rely heavily on lichens in winter.
Dall sheep Ovis dalli In parts of Alaska and Yukon, overlaps with caribou on alpine and subalpine ranges. Both are migratory or seasonally elevational in some populations, and both face similar constraints from snow and ice conditions and similar predator guilds (wolves, bears, and eagles preying on young).
Saiga antelope
Saiga antelope Saiga tatarica Functional analogue on open steppe: a highly mobile grazing/browsing ungulate that forms large migratory aggregations and tracks seasonal forage across vast landscapes, exhibiting movement ecology similar to migratory barren-ground caribou (niche similarity despite occurring in different regions).
Pronghorn
Pronghorn Antilocapra americana Ecological analogue in North America as a long-distance migrant occupying open habitats. Like migratory caribou, its movements are tightly linked to seasonal plant phenology, snow conditions, and avoidance of the energetic costs of deep snow.
Arctic hare
Arctic hare Lepus arcticus Shares Arctic tundra environments and winter snow-foraging constraints. Both are prey for some of the same predators (e.g., wolves opportunistically; eagles primarily target leverets/calves) and depend on seasonal shifts in available tundra vegetation.

Caribou, which are sometimes referred to as reindeer, are notable for their antlers, growing back bigger every single year. They have unique stomach bacteria that allow them to eat many different types of food. The name caribou covers several different species, but hunting has led to the extinction of two of them.

5 Incredible Facts!

Here are a few fun facts about the Caribou.

  • Both males and females have antlers.
  • The stomach of a caribou has four chambers.
  • The color of caribou fur largely depends on where they live. Lighter colors are associated with northern regions, while caribou with dark coats live in southern areas instead.
  • The scientific name of the caribou translates to “reindeer reindeer.”
  • Caribou are often hunted for their meat.

You can click this link for more incredible facts about Caribou.

Scientific Name

The caribou, which goes by the scientific nameRangifer tarandus,” is part of the Mammal class and Cervidae family. Caribou go by several names, including reindeer, qalipu, and tuktu.

The name “Rangifer tarandus” was chosen by Carl Linnaeus, describing the reindeer genus as “Rangifer.” It is Latin, and it literally means “reindeer.” Tarandus, however, comes from a New Latin translation of the ancient word “tárandos,” which means reindeer as well. This species is called reindeer only in Europe/Asia and among semi-domesticated animals in North America. Otherwise, they are caribou.

History and Species

Although the reindeer is famously associated with the Baltic and Scandinavian countries and with the Siberian and Mongolian regions, the caribou actually evolved in North America and spread to Eurasia. Fossils of caribou in Alaska and the Yukon can be dated to 1.3–1.8 million years ago.All caribou/reindeer throughout the world are considered to be the same species, but there are 7 subspecies:

  • Barrenground (Rangifer tarandus granti) – the main type in Alaska, also in Canada
  • Svalbard (R.t platyrhynchus),
  • European (R.t. tarandus),
  • Finnish forest reindeer (R.t. fennicus),
  • Greenland (R.t. groenlandicus),
  • Woodland (R.t. caribou) – one small herd in Alaska, also in Canada
  • Peary (R.t. pearyi) – in Canada

Appearance

The caribou, better known as a reindeer, has massive antlers that start right at the brow. Both females and males have antlers, and they are the only cervid species to do so. The antlers begin to grow in March or April for males, but the females will start their growth in May and June. The antlers feature a velvety texture that is deep through to blend with the rest of their body. The rack can ultimately reach a height of 3 feet tall.

Among deer species, caribou hooves are unique. They are cloven with four toes, two of which are called dew claws, that spread widely to provide support. This structure helps the caribou to walk easily on many surfaces, including soft tundra and snow. Caribous hooves are the widest and roundest of any deer, and they actually change size according to the season. The feet also function as paddles when caribou swim, and by the way, they are excellent swimmers.

The color of the fur of this mammal largely depends on the season, the subspecies, and the individual caribou animal. In the north, the fur tends to be white to blend with the snow, which is typical of the Peary caribou. Darker fur is seen in southern areas, and the darkest caribou of all tends to be the boreal woodland caribou.

Caribou are the only deer species to have hair covering from head to toe, even covering the nose and growing under the feet to help give traction. Their fur is actually in two layers for better warmth and protection.

Male Caribou grazing on Toklat river basin.

Male Caribou average 350-400 pounds, but the larger species can get up to 700 lbs..

The species will also determine how large the animal is. Females are usually smaller than males, starting at 64 inches in length, 33 inches in height, and about 180-260 lbs. in weight. The smallest reindeer comes from Svalbard and has a height of just 31 inches. Males will vary more among the species, starting at 350 lbs. However, the biggest species have males that can be well over 700 lbs. That’s slightly less than the weight of an Arabian riding horse.

Behavior

The social behavior of the Caribou largely depends on the locations and species. Reindeer occur in both migratory and sedentary populations, and their herd sizes vary greatly in different regions.

Those caribou that migrate will cover about 1,500 miles annually with impressive speed as they search for food. During migration, there’s a chance that the animal can become susceptible to parasites, making them incredibly weak. Parasites can vary from one species to the next, but migration can cause them to lose their weakest caribou during travel.

Caribou can run up to 50 mph (80 kmh). Even the baby caribou can outrun an Olympic runner when they’re barely 24-hours old. Herds that migrate during the springtime could have up to 500,000 caribou animals, though autumn migrations are much smaller.

Reindeer are the only type of deer that have been partially domesticated. Some cultures use them as work animals (like pulling a sleigh for Santa Claus!) They are, of course, an important source of food clothing, and shelter among Arctic natives .

Habitat

Caribou previously were found only in certain locations in Scandinavia, Mongolia, Siberia, northeastern Europe, Greenland, Russia, and some parts of China. However, they can also be found in North America as low as Maine and as high as Alaska, and in most northern areas of Canada. Animals that live further north tend to have a white coat that conceals them in the snow, while southern locations feature a dark brown color that shields them in the woods.

These mammals can live in the tundra or the forest, though migration and climate change can be credited for their lower numbers across large areas.

Predators and Diet

What eats Caribou?

Although the caribou is a rather large animal, they still have many predators. In some areas, the caribou is hunted for its meat by humans. In fact, this is part of the reason that caribou disappear in some locations.

The calves are at the greatest risk of predators since golden eagles and wolverines tend to go after them. However, polar bears and brown bears will go after reindeer of all sizes. They are also extremely at risk of bloodsucking insects like mosquitoes, black flies, and the reindeer nose botfly.

What do Caribou eat?

Caribou fall under a unique category of ruminants, meaning that their stomach has a total of four chambers. They are the only large animal that can use lichen to make glucose. However, their primary diet consists of willows, birches, grasses, and other plant life. They’ll even eat the antlers of other caribou before they are shed to get supplementary nutrients. Some will even eat mushrooms.

To find plants in the snow, caribou are equipped with hooves that are hollowed out so that they can be used like scoops to dig for food.

Caribou are not really carnivorous unless they have to be. However, if their springtime nourishment isn’t quite as plentiful as they’d like, they’ll look for what’s available (like rodents, fish, and eggs).

Reproduction, Babies, and Lifespan

During the fall (from late September to early November), reindeers will mate. The males will battle each other to win the females, locking their antlers as they push against each other.This is called being in rut or performing ruts.

The victorious males will likely get as many as 20 females to mate with, but they will lose a lot of their weight in the process. Caribou have a gestation period of 222-234 days (about 7-8 months).

Females generally take on only one mate at this time, siding with the more dominant males. They are incredibly careful while choosing where they will give birth, looking for an area that generally doesn’t have many predators. The herd follows the doe as she looks for the right place to give birth to a single calf, which typically weighs about 13 lbs.

Babies are called calves, and they will be running in less than two hours after birth with great speed. They learn to forage as early as 45 days, although they will nurse from their mothers until the autumn when they are about 3-4 months old.

Females live longer than males, but the average lifespan of a caribou is 10-15 years.

Caribou Population

Since Caribou consist of many different species, there are a few subspecies that have already gone extinct. As recently as 2015, reindeer were on the Vulnerable list with the IUCN since they have seen a decline of 40% in the last 2.5 decades. However, the massive population of the caribou specifically ensures that the main species is not at any risk right now.
The Rangifer tarandus dawsoni already went extinct, and the Rangifer tarandus pearyi is one of the endangered species.

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Sources

  1. USGS / Accessed August 28, 2021
  2. Wikipedia / Accessed August 28, 2021
  3. Animal Corner / Accessed August 28, 2021
  4. Alaska Department of Fish and Game / Accessed August 28, 2021
  5. FDA / Accessed August 28, 2021
  6. Britannica / Accessed August 28, 2021

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Caribou FAQs (Frequently Asked Questions)

Caribou and Moose are often compared to one another since they can both be found in Arctic environments. In general, moose are larger while Caribou generally have lighter coats and “thinner” horns that have a very different structure.