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Species Profile

Natterjack

Epidalea calamita

The stripe-backed toad that runs and roars
iStock.com/kwasny221

Natterjack Distribution

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Natterjack toad sitting at the water's edge.

At a Glance

Wild Species
Also Known As Natterjack, European natterjack, Yellow-banded toad, sapo corredor (Spanish), crapaud calamite (French)
Activity Nocturnal+
Lifespan 6 years
Weight 0.08 lbs
Status Least Concern
Did You Know?

Adults are typically 60-70 mm snout-vent length (rarely to ~80 mm), smaller than the common toad (Bufo bufo).

Scientific Classification

The natterjack toad is a small to medium-sized true toad native to parts of western and central Europe, noted for its distinctive loud, rasping call and preference for sandy, open habitats.

Kingdom
Animalia
Phylum
Chordata
Class
Amphibia
Order
Anura
Family
Bufonidae
Genus
Epidalea
Species
calamita

Distinguishing Features

  • Often shows a yellow stripe running down the midline of the back (a common field mark, though not always prominent)
  • Shorter legs and a characteristic running gait (rather than long hops)
  • Very loud, harsh, sustained male call during breeding season
  • Prefers sandy, open habitats compared with many other European toads

Physical Measurements

Males and females differ in size

Length
♂ 2 in (2 in – 3 in)
Weight
♂ 0 lbs (0 lbs – 0 lbs)
♀ 0 lbs (0 lbs – 0 lbs)
Top Speed
2 mph
running
Poisonous

Appearance

Primary Colors
Secondary Colors
Skin Type Dry, granular-warty skin with prominent parotoid glands; adapted for terrestrial life in open sandy habitats.
Distinctive Features
  • Adult snout-vent length typically 6-8 cm; females average larger than males (Arnold & Ovenden, 2002).
  • Compared with Bufo bufo, generally smaller and more lightly built; favors open sandy heaths/dunes rather than woodland (Beebee & Griffiths, 2000).
  • Often shows a pale/yellow mid-dorsal stripe, but stripe intensity varies and can be weak or absent (Arnold & Ovenden, 2002).
  • Parotoid glands evident behind eyes; warts often with darker tips giving a peppered look (Beebee & Griffiths, 2000).
  • Breeds in very shallow, warm, temporary pools; eggs laid in long strings typical of true toads (Beebee & Griffiths, 2000).
  • Males produce a loud, rasping call during breeding choruses, a key field cue in open habitats (Beebee & Griffiths, 2000).
  • Conservation: listed as Least Concern globally (IUCN), but protected/priority in parts of its European range where habitats are fragmented (e.g., regional/national protections).

Sexual Dimorphism

Sexes are similar in color and pattern, but females are typically larger. Breeding males develop nuptial pads and a conspicuous subgular vocal sac used for the species' loud rasping call; females lack these features.

♂
  • Typically smaller body size than females (Arnold & Ovenden, 2002).
  • Darkened subgular vocal sac when calling in breeding season.
  • Nuptial pads on inner fingers during breeding, aiding amplexus.
♀
  • Typically larger, more robust body (Arnold & Ovenden, 2002).
  • No vocal sac; does not produce the rasping advertisement call.
  • Lacks nuptial pads; abdomen may appear fuller when gravid.

Did You Know?

Adults are typically 60-70 mm snout-vent length (rarely to ~80 mm), smaller than the common toad (Bufo bufo).

A narrow yellow dorsal stripe is common and is a key field mark in much of its range (though it can be faint/absent in some individuals).

Males give a loud, rasping "ratchet-like" call that can carry well over 1 km on still nights-one of the loudest amphibian calls in Europe relative to body size.

Unlike many toads that mainly hop, natterjacks often RUN quickly across open sand on relatively short legs.

Breeding is concentrated in very shallow, sun-warmed, temporary pools; faster tadpole development helps them beat pond drying.

Females lay eggs in paired strings; a single clutch commonly contains thousands of eggs (often cited ~2,000-7,000).

Longevity can reach about 15 years in the wild under good conditions (shorter is common where predation and drought are high).

Unique Adaptations

  • Open-sand specialist: favors sparsely vegetated dunes, coastal systems, and lowland heaths-habitats avoided by many other European toads, including Bufo bufo.
  • Warm-water breeding strategy: preference for shallow, sunlit pools accelerates embryo and tadpole growth compared with cooler, deeper ponds.
  • Chemical defense: parotoid glands secrete bufotoxins (a hallmark of true toads, Bufonidae) that deter many predators.
  • Camouflage for sand and heath: mottled tan/olive coloration blends with dunes and bare ground; the pale/yellow dorsal stripe can break up the body outline.
  • Tolerance of challenging coastal conditions: populations often persist in windy, salty, and shifting dune environments where few amphibians thrive (notably in coastal dune systems).

Interesting Behaviors

  • Nocturnal foraging in open habitats: emerges at dusk to hunt beetles, ants, spiders, and other invertebrates on bare sand and short turf.
  • Rapid terrestrial locomotion: frequently runs in bursts rather than making long hops-useful in windswept dunes and sparsely vegetated heaths.
  • Explosive chorus calling: males call from very shallow water (often only a few centimeters deep), forming loud choruses that advertise breeding sites across large areas.
  • Opportunistic breeding in ephemeral pools: uses newly formed rain pools, dune slacks, and wheel-rut puddles where fish are absent and water warms quickly.
  • Burrowing and concealment: shelters by day in self-made scrapes or shallow burrows in sand; also uses mammal burrows and vegetation clumps for cover.
  • Strong site fidelity: adults often return to the same breeding area year after year when habitat remains suitable.
  • Seasonal activity shaped by weather: surface activity increases in warm, humid conditions; during drought or cold snaps, individuals stay buried to reduce water loss.

Cultural Significance

The natterjack toad (Epidalea calamita) is a symbol for saving coastal dunes and lowland heath. IUCN lists it as Least Concern, but it is rare in some countries. It is protected by the EU Habitats Directive and laws (e.g., Great Britain). Its 'nattering' call marks healthy dune slacks and pools.

Myths & Legends

Natterjack toad (Epidalea calamita) was often linked in Medieval and early modern Europe to witchcraft; dune and heath toads were called witches' helpers or used in charms and potions, shaping witch stories.

In Britain people believed a 'toadstone'—a jewel from a toad used as an amulet against poison and sickness. Not tied to one species, it is part of stories about native toads like natterjack (Epidalea calamita).

In parts of Europe, folk sayings said the Natterjack toad (Epidalea calamita) near a home was an omen about weather, luck, or household protection, linked to its secret, night habits and sudden appearance after rain.

Name lore: The English name "natterjack" links to its nattering call and the 'jack' everyman suffix; this helped make the natterjack toad (Epidalea calamita) the loud toad of British sands and heaths.

In heath and dune stories, people called the loud rasping calls of the natterjack toad from shallow pools the 'singing' of the land after warm rain—a sign the dunes and heath were alive and renewed.

Conservation Status

LC Least Concern

Widespread and abundant in the wild.

Population Decreasing

Protected Under

  • EU Habitats Directive 92/43/EEC (Annex II and Annex IV)
  • Bern Convention on the Conservation of European Wildlife and Natural Habitats (Appendix II)
  • UK Wildlife and Countryside Act 1981 (as amended) / Conservation of Habitats and Species Regulations (where applicable in range states)

Life Cycle

Birth 3000 tadpoles
Lifespan 6 years

Lifespan

In the Wild
3–15 years
In Captivity
5–20 years

Reproduction

Mating System Promiscuity
Social Structure Aggregation Group
Breeding Pattern Transient
Fertilization Broadcast Spawning
Birth Type Substrate_spawning

In spring-early summer, males form loud choruses at shallow ponds and may amplex multiple females; females typically mate once per spawning. External fertilization occurs as 1,500-4,000 eggs are released in strings; no pair bond or parental care.

Behavior & Ecology

Social Breeding chorus Group: 20
Activity Nocturnal, Crepuscular, Diurnal
Diet Insectivore Ants (Formicidae) - frequently reported as a major dietary component in sandy/heathland populations
Seasonal Migratory, Hibernates 2 mi

Temperament

Generally non-aggressive; relies on crypsis, rapid running, and short hops when disturbed.
Breeding males show strong calling-site fidelity within ponds; interactions are primarily acoustic.
Outside breeding, adults are mostly solitary and dispersive between feeding areas and refuges.
Breeding aggregation size varies widely with pond hydroperiod and local population density.
Philopatry to traditional breeding ponds is common; dispersal increases after pond loss.
Longevity reported up to ~15 years in the wild (e.g., Beebee & Griffiths, 2000).

Communication

Male advertisement call: loud, rasping/trilling call; carries long distances Often cited to ~1-2 km) (Beebee & Griffiths, 2000
Release call: brief call given when clasped by another toad Typical anuran behavior
Distress call: short squeak when handled or attacked Reported in many bufonids
Tactile signaling during amplexus and positioning while spawning Clasping, body contact
Visual cueing via inflated vocal sac and calling posture at pond margins Mate attraction
Chemical cues likely aid mate/pond recognition at close range Documented broadly in amphibians
Call intensity and chorus structure vary with temperature, wind, and male density.

Habitat

Biomes:
Temperate Grassland Temperate Forest Mediterranean Freshwater Wetland
Terrain:
Coastal Plains Hilly Valley Island Sandy Riverine +1
Elevation: Up to 7874 ft

Ecological Role

Nocturnal terrestrial mesopredator of invertebrates in dune, heath, and other open sandy ecosystems; also an important prey item for higher predators.

Regulates populations of ground-dwelling insects and other invertebrates (including ants, beetles, flies) Transfers energy from invertebrate communities to higher trophic levels (birds, reptiles, mammals) Links aquatic and terrestrial food webs via larval-to-adult metamorphosis (export of aquatic productivity to land)

Diet Details

Human Interaction

Domestication Status

Wild

Natterjack toad (Epidalea calamita) is not domesticated and has no history of breeding for pets, food, or work. People mainly interact through conservation, habitat restoration, monitoring, and rare licensed captive breeding to release them back into the wild. They breed in warm shallow pools, have very loud calls, and are legally protected in much of Europe.

Danger Level

Low
  • Skin secretions contain bufotoxins typical of bufonid toads; can irritate eyes/mucous membranes and may cause nausea if ingested (primary risk is to pets that mouth/biting toads).
  • Handling amphibians (including this species) can pose a low but non-zero zoonotic risk (e.g., Salmonella transmission) if hygiene is poor; wash hands after contact.
  • Indirect/public-nuisance interaction: loud breeding choruses near human developments can lead to complaints, though this is not a physical danger.

As a Pet

Not Suitable as Pet

Legality: Natterjack toad (Epidalea calamita) is mostly illegal to take or keep from the wild without permits. Many countries protect it, so pets are rare and only allowed for licensed institutions or approved keepers.

Care Level: Expert Only

Purchase Cost:
Lifetime Cost: $800 - $2,500

Economic Value

Uses:
Ecosystem service: insect predation (consumes a wide range of terrestrial invertebrates; locally valued for natural pest control) Conservation value/flagship species (habitat restoration and protected-area management for dunes, heathlands, and ephemeral pond systems) Education and research value (used in ecological, behavioral, and conservation biology studies; public engagement via nature reserves) Controls pest population (insectivorous predator)
Products:
  • No major commercial products; value is primarily ecological (pest control, biodiversity) and informational (education/research outputs).

Relationships

Predators 8

Grass snake
Grass snake Natrix natrix
European adder
European adder Vipera berus
European hedgehog Erinaceus europaeus
Red fox
Red fox Vulpes vulpes
European badger Meles meles
Grey heron
Grey heron Ardea cinerea
Black-headed gull Chroicocephalus ridibundus
Carrion crow Corvus corone

Related Species 5

Common toad
Common toad Bufo bufo Shared Family
Spined toad Bufo spinosus Shared Family
European green toad Bufotes viridis Shared Family
Balearic green toad Bufotes balearicus Shared Family
Eastern green toad Bufotes sitibundus Shared Family

Ecological Equivalents 4

Animals that fill a similar ecological role in their ecosystem

Common spadefoot toad Pelobates fuscus Natterjack toad (Epidalea calamita) and spadefoot toads overlap in parts of Europe: both are nocturnal, burrow in loose sandy soil, and breed in temporary warm, fish-free pools often formed after rain.
Common frog
Common frog Rana temporaria Often found together and sharing breeding ponds; both the natterjack and Rana temporaria have aquatic tadpoles and terrestrial adults. The natterjack prefers open sandy heaths and dunes, while Rana temporaria uses cooler, more vegetated ponds.
Moor frog Rana arvalis Where their ranges meet, the natterjack toad and the moor frog both use open lands with shallow wetlands and breed in small, sunlit waters. They overlap most in lowland open habitats; the moor frog prefers marshy, peaty areas.
Parsley frog Pelodytes punctatus In western and southwestern Europe it can co-occur in sandy or open habitats and use small temporary ponds. Both species are nocturnal foragers on small invertebrates and use warm, fishless breeding sites, although Pelodytes is generally smaller and has a more Mediterranean-leaning distribution.
The natterjack is a species of toad found in coastal areas of Europe with a distinctive warty skin featuring a mix of colors such as green, brown, and cream.
The natterjack is a species of toad found in coastal areas of Europe with a distinctive warty skin featuring a mix of colors such as green, brown, and cream.

“A natterjack toad can lay up to 7500 eggs in a single clutch”

The Natterjack toad is native to Europe. It looks a lot like a common toad with the exception of a vertical yellow line on its back. It’s carnivores eating woodlice, worms, and spiders. These amphibians are active at night looking for prey. Their lifespan is 15 to 20 years.

5 Incredible Natterjack Facts!

Natterjack toad sitting at the water's edge.

The rapid movement style of crawling, instead of hopping, has earned this amphibian the moniker “running toad”.

  • This amphibian is known as the running toad because of the speedy way it crawls instead of hops
  • The sound of its loud call can travel up to three miles
  • They have a lifespan going up to 20 years
  • They live in a habitat with marshes and sand dunes
  • This amphibian display the same colors as a common toad but with a yellow stripe down its back

Scientific Name

Male Natterjack toad singing to attract females.

The scientific name of the natterjack toad is Epidalea calamita.

The natterjack toad’s scientific name is Epidalea calamita. The word Epidalea is the genus of this toad while the meaning of the word calamita is magnetic. This refers to the belief that the moon and its magnetic pull draws, European toads (like magnets) to shallow ponds during mating season.

The nickname of this amphibian is the running toad because of the way it moves. This nickname doesn’t have a literal meaning behind it. Though this amphibian doesn’t actually run, it moves so swiftly that it looks like it’s running! It’s a member of the Bufonidae family and is in the Amphibia class.

Evolution and Origins

The natterjack toad is the singular type of toad that is indigenous to Ireland. It can be located in two counties, namely County Kerry (Dingle Peninsula and Derrynane) and County Wexford, where it was introduced to a dune site.

However, in mainland Europe, specifically in the southern areas of its distribution range, it resides inland, occupying a range of habitats.

Bufo calamita, or the Natterjack toad, is commonly used in European traditional medicines and rituals, with its significance extending from Germany to East Anglia.

Additionally, it is unique among the toad species as it is native to Ireland, which has a rich history of folklore and magic practices that have been a source of inspiration for the British Isles.

Appearance and Behavior

Natterjack toad sitting on grass.

An adult natterjack has wart-covered skin that displays a combination of hues such as green, brown, and cream.

The wart-covered skin of an adult natterjack features a mixture of colors including green, brown, and cream. The identification of this amphibian is easily done by looking for the yellow stripe down its back. This is one of its most obvious features. Its back legs are also shorter than a common toad’s hind legs.

At two to three inches long, natterjacks are smaller in size than common toads. A three-inch natterjack is equal in length to one-and-a-half golf tees. Alternatively, common toads can grow to be six inches in size. The heaviest a natterjack can weigh is a little over half an ounce.

Though this amphibian is just three inches long at the most, it has some defenses against predators. For one, its dark colors allow it to blend well into its marshy habitat. Also, it can puff up its body and stretch out its legs to make itself appear larger.

This can be enough to scare some predators away. In addition, a natterjack has glands on its head that release a milky poison. When grabbed by a predator like a hedgehog or a heron, the natterjack releases poison causing the predator to let go. Once it’s on the ground, the amphibian can get away quickly by moving in its unique crawling gait.

Another defense of this amphibian is it is active at night. Moving under cover of darkness can help this toad to remain hidden and safe from all of the predators active during the day.

Natterjacks are solitary with the exception of the breeding season when these amphibians gather at shallow ponds. A group of toads is called a knot. Breeding season is when the distinctive male mating call can be heard for up to three miles.

Some people think their call sounds like an ‘er, er’ sound. If you watch a male natterjack call to females, its neck expands into a bubble to send the call even further into the distance. This is one of those facts you almost have to see to believe. Furthermore, you’d have to ask a male natterjack for the exact meaning behind its strange-sounding call.

Generally speaking, these amphibians have a shy nature, hiding in marsh grass and under rocks until it’s time to go out and hunt for food at night. They aren’t aggressive creatures.

Habitat

Natterjack Toad eating earthworms.

The natterjack toad is a type of European toad that can be found in various coastal regions including Spain, England, France, Ireland, and Germany, among others.

The natterjack is a European toad. It lives in the coastal areas of Spain, England, France, Ireland, and Germany, to name a few. Their habitat includes sand dunes, marshes, and wetlands. They gather in shallow pools of warm water during mating season. This is where they mate and where females lay their strings of eggs.

If you were looking for this amphibian in a marshy area, you’d probably find some hiding under rocks because these creatures like to dig in the soft sand. Or, you may see one or two sticking their heads up out of the marsh water. However, these tiny creatures blend in so well with the dark water and vegetation around them, you could be looking at one and not even realize it!

These amphibians are nocturnal, so they are out hunting and calling each other at night. They have big eyes and thin pupils giving them good vision after the sunlight disappears. They need good vision to capture the insects they need to nourish themselves.

Predators and Threats

The life cycle of a natterjack has several stages including egg, tadpole, juvenile, and adult. A natterjack in the tadpole stage eats aquatic vegetation while an adult is a carnivore eating insects of many types.

What does a natterjack toad eat?

Natterjacks eat spiders, woodlice, worms, snails, and other small insects. These amphibians hide beneath marshy plants waiting to capture insects passing by.

What eats natterjack toads?

Herons, hedgehogs, and foxes are some of the predators of this tiny amphibian. These animals share the same habitat as the running toad and look for food in or near the water. These predators are likely to consume this amphibian’s tadpoles and eggs as well.

In terms of conservation, natterjacks are listed as Least Concern but with a decreasing population. Their numbers are dropping in part due to habitat loss. Their habitat is dwindling as a result of farmland expansion and natural changes in the landscape. They are also at risk of a deadly disease called chytridiomycosis. Chytridiomycosis travels quickly among groups of amphibians. This disease is studied by examining the tissue of a dead amphibian.

Reproduction and Life Cycle

Closeup on a small juvenile Natterjack Toad, a rare and protected species crawling on sandy soil.

This toad typically starts its breeding season in March or April and continues it throughout the summer.

The breeding season of this toad begins in March or April and continues into the summer. Males form large groups or knots, in and around shallow ponds. From there, they call for female mates by making a loud, raspy sound. These amphibians have several partners during the breeding season.

Female toads lay eggs in warm shallow water. This is another one of those facts to help you with identification. While common toads lay two strings of eggs, a natterjack toad lays just one. But, that one string can have up to 7500 eggs! Once the eggs are laid the female leaves her young. It takes eight to fifteen days for the eggs to hatch moving this toad to the next step in its life cycle.

As a note, male natterjacks have been known to sit at the edge of shallow pools where eggs are laid. Some biologists believe these males are standing guard over the eggs and tadpoles. If a smaller predator comes along, a male puffs up its body in an effort to scare the intruder away from the vulnerable eggs.

Natterjack toad babies called tadpoles are less than an inch in size. One of the reasons this amphibian lays so many eggs is there’s a good chance lots of the eggs or tadpoles will be eaten by fish, frogs, and other predators. The more tadpoles there are, the greater the chance some will survive long enough to grow into adult natterjacks.

Tadpoles are less vulnerable to passing predators than the eggs of this amphibian. Tadpoles are able to hide among the floating vegetation in a shallow pond and consume aquatic plants for nourishment.

The life cycle of this toad from egg to juvenile takes about eight weeks. The lifespan of this amphibian is 15 to 20 years.

Population

The population of this amphibian is unknown. It’s listed as Least Concern but is decreasing in number. In the world of European toads, it’s considered rare.


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Sources

  1. Wikipedia / Accessed February 11, 2022
  2. Amphibian and Reptile Conservation / Accessed February 11, 2022
  3. Cornell Wildlife Health Lab / Accessed February 11, 2022
  4. Wikipedia / Accessed February 11, 2022
  5. IUCN Redlist / Accessed February 11, 2022
  6. Let's Go Wild / Accessed February 11, 2022
Rebecca Bales

About the Author

Rebecca Bales

Rebecca is an experienced Professional Freelancer with nearly a decade of expertise in writing SEO Content, Digital Illustrations, and Graphic Design. When not engrossed in her creative endeavors, Rebecca dedicates her time to cycling and filming her nature adventures. When not focused on her passion for creating and crafting optimized materials, she harbors a deep fascination and love for cats, jumping spiders, and pet rats.
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Natterjack FAQs (Frequently Asked Questions)

This amphibian in its adult stage eats a diet of spiders, beetles, woodlice, snails, worms, and sometimes dragonflies.

During the tadpole stage of its life cycle, they nibble on aquatic plants in their habitat. So, these amphibians have a different type of diet in various stages of its life cycle. This is true with many types of amphibians.