S
Species Profile

Sardines

Clupeidae

Small fish, huge impact
Dennis Forster/Shutterstock.com

Sardines Distribution

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Invasive Species

This map shows coastal regions where Sardines are found.

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Found in 104 countries

School of sardines swimming from left to right

At a Glance

Family Overview This page covers the Sardines family as a group. Stats below are general traits shared across the family.
Also Known As Pilchard, Brisling, Sprat
Diet Filter Feeder
Activity Diurnal+
Lifespan 6 years
Weight 0.25 lbs
Status Not Evaluated
Did You Know?

"Sardine" is a market name: many Clupeidae species (pilchards, sprats, young herrings) can be sold as sardines depending on region and size.

Scientific Classification

Family Overview "Sardines" is not a single species but represents an entire family containing multiple species.

Sardines are small, silvery, schooling pelagic fishes best represented by the herring family (Clupeidae). They are plankton-feeders and are ecologically important forage fish, as well as a major global fishery and canned-food product.

Kingdom
Animalia
Phylum
Chordata
Class
Actinopterygii
Order
Clupeiformes
Family
Clupeidae

Distinguishing Features

  • Small to medium-sized, laterally compressed, silvery fishes
  • Single soft-rayed dorsal fin; generally no fin spines
  • Forked tail; schooling behavior
  • Filter-feeding on plankton using gill rakers
  • Key prey for larger fish, seabirds, and marine mammals

Physical Measurements

Males and females differ in size

Length
♂ 10 in (2 in – 1 ft 4 in)
♀ 8 in (2 in – 1 ft 3 in)
Weight
♂ 0 lbs (0 lbs – 1 lbs)
♀ 0 lbs (0 lbs – 0 lbs)
Top Speed
19 mph
swimming

Appearance

Primary Colors
Secondary Colors
Skin Type Thin, delicate cycloid scales with heavy guanine deposition producing a silvery sheen; typically coated in mucus; many species have a ventral keel with scutes (bony belly plates).
Distinctive Features
  • Size range across Clupeidae: ~3-60 cm total length (most commonly ~10-30 cm).
  • Mass range: a few grams in smallest species to ~2-3 kg in largest shads/hilsas.
  • Typical body form: laterally compressed, streamlined, and deep-bodied to moderately slender.
  • Single mid-body dorsal fin; forked caudal fin adapted for sustained schooling swimming.
  • Large eyes; small, terminal to slightly upturned mouth; gill rakers often well-developed for plankton filtering.
  • Ventral keel with scutes (sharp-edged belly plates) common; extent varies among genera.
  • Often a dark shoulder spot behind the gill cover; additional flank spots may occur in some lineages.
  • Lifespan range across family: commonly ~1-10 years, with some species reaching ~12-15 years.
  • Behavior/ecology (general): strongly schooling pelagic or coastal; many perform diel vertical migrations.
  • Feeding (general): primarily plankton-feeders (filtering phytoplankton/zooplankton), but some shift to larger prey as they grow.
  • Reproduction (general): broadcast spawners producing many small pelagic eggs; timing varies regionally and by species.
  • Habitat variation: many marine coastal/pelagic; some estuarine; several anadromous or primarily freshwater species.
  • Ecological role: key forage fish linking plankton to predators (seabirds, marine mammals, larger fishes).
  • Commercial context: major global fisheries and canned products; population size and body condition can vary strongly with ocean productivity and fishing pressure.

Sexual Dimorphism

Sexual dimorphism is usually subtle. Females are often slightly larger and deeper-bodied when gravid, while males in some species develop seasonal rough scales or small breeding tubercles and may be slimmer outside spawning periods.

♂
  • Often slightly slimmer body profile outside spawning periods.
  • Seasonal breeding tubercles or roughened scales reported in some clupeids.
  • May mature at smaller sizes than females in certain populations.
♀
  • Often slightly larger on average in many species/populations.
  • Noticeably rounder abdomen when carrying roe during spawning season.
  • May show higher body mass at the same length when gravid.

Did You Know?

"Sardine" is a market name: many Clupeidae species (pilchards, sprats, young herrings) can be sold as sardines depending on region and size.

Family size spans roughly 2-75 cm, from tiny sprat-like clupeids to large shads.

Lifespans vary widely: fast-growing small sardines may live ~1-5 years, while some herrings/shads can reach ~15-25 years.

Many clupeids are filter-feeders, using long gill rakers to strain plankton-turning microscopic productivity into food for bigger animals.

Several shads (genus Alosa, in Clupeidae) can detect ultrasound-an unusual fish sense thought to help avoid echolocating dolphins.

Their tight schools are a defense strategy and a feeding tool, but also make populations sensitive to changes in ocean conditions and fishing pressure.

Clupeids fuel famous wildlife spectacles, including "sardine run"-style events where predators concentrate around dense baitfish schools.

Unique Adaptations

  • Silvery, reflective sides (countershading): reduces visibility in open water by mirroring the environment-effective across many pelagic clupeids.
  • Gill rakers adapted for plankton: comb-like structures that enable efficient filtering; raker spacing differs among species and influences prey size.
  • Abdominal scutes: a keel of sharp, bony belly plates common in Clupeidae that may aid hydrodynamics and/or predator deterrence.
  • Otophysic connection (hearing enhancement): many clupeids have special links between the swim bladder and inner ear that improve sound detection in pelagic habitats.
  • Rapid growth and early maturity (common but variable): especially in small sardine-like species, supporting quick population rebounds when conditions are favorable.

Interesting Behaviors

  • Schooling at multiple scales: individuals form coordinated groups that can merge into massive aggregations; school size and cohesion vary by species, age, and predator pressure.
  • Diel movement patterns: many species shift depth between day and night (often deeper by day, shallower at night), though patterns vary with water clarity and food.
  • Plankton feeding strategies: some primarily filter-feed on tiny plankton, while others switch to picking larger prey (e.g., copepods) when available.
  • Spawning synchrony: many are broadcast spawners releasing eggs and sperm in the water column; timing can be tightly linked to temperature, currents, and plankton blooms.
  • Long-distance migrations (variable): some are coastal pelagic movers; others (notably shads and river herrings) are anadromous-migrating from the sea into rivers to spawn.
  • Boom-bust dynamics: recruitment can swing strongly year to year with ocean conditions, creating natural population variability that complicates fisheries management.

Cultural Significance

Clupeidae sardines are key forage fish that support seabirds, marine mammals, and large fish, and sustain human jobs, canned-fish markets, and coastal foods. They are cheap, rich in omega-3 fats, but their numbers swing with climate, raising worries about overfishing and ecosystem-based management.

Myths & Legends

Name-and-place tradition: "Sardine" is widely linked to Sardinia (Mediterranean), reflecting an old association between these fish and the island's waters and trade routes.

Japan (spring festival folk practice): grilled sardine heads stuck on holly branches are traditionally placed at entrances to ward off demons during seasonal rites.

Cornwall (pilchard lore): in Cornish tradition, pilchards (often classed as sardines in commerce) are tied to local saints and coastal blessings, reflecting the fish's historical role in survival and prosperity.

Mediterranean religious and festival associations: in Iberian and Mediterranean coastal towns, sardines became emblematic festival foods (especially summer saint-day celebrations), blending maritime identity with communal ritual meals.

Conservation Status

NE Not Evaluated (family-level). Across Clupeidae, individual species range from Least Concern to threatened categories (including VU/EN in some restricted-range, migratory river/estuary-spawning shads and localized endemics), with many also listed as DD/NE. Measurements vary widely: smallest members ~3-5 cm total length; largest herrings/shads reach ~60-70+ cm. Lifespan spans roughly ~1-3 years in many small, fast-growing sardine-like species to ~10-25+ years in larger, slower-growing clupeids. Ecology/behavior generalizations: typically schooling, pelagic-to-neritic planktivores that form major forage-fish biomass; many are highly migratory and environmentally driven (strong recruitment variability), while others are coastal residents or anadromous/estuarine spawners-conservation risks differ strongly by habitat and fishery exposure.

Has not yet been evaluated against the criteria.

Population Unknown

Protected Under

  • National and regional fisheries laws setting catch/effort controls, seasonal closures, and gear rules (varies by country)
  • EU Common Fisheries Policy (CFP) with TACs/management plans for relevant small pelagic stocks
  • United States Magnuson-Stevens Fishery Conservation and Management Act (stock assessments and harvest control rules for managed clupeids)
  • Regional Fisheries Management Organizations (RFMOs) and scientific advisory bodies where applicable (stock-based management in shared waters)
  • Marine Protected Areas and coastal habitat protections that incidentally safeguard spawning/nursery zones (coverage varies widely)

You might be looking for:

European pilchard (European sardine)

32%

Sardina pilchardus

Often considered the ‘true sardine’ in Europe and a common canned sardine species in the Atlantic/Mediterranean.

Pacific sardine

26%

Sardinops sagax

Major sardine species in the Pacific (e.g., off North and South America, Australia/NZ).

Japanese sardine

16%

Sardinops melanostictus

Important commercial ‘sardine’ in the Northwest Pacific, especially around Japan.

Round sardinella

14%

Sardinella aurita

Commonly marketed as sardines in parts of the Atlantic and Mediterranean; warm-water clupeid.

European sprat (sometimes sold as ‘brisling sardines’)

12%

Sprattus sprattus

A clupeid often canned and marketed as small sardines in some regions.

Life Cycle

Birth 50000 frys
Lifespan 6 years

Lifespan

In the Wild
1–25 years
In Captivity
0.25–10 years

Reproduction

Mating System Promiscuity
Social Structure Aggregation Group
Breeding Pattern Not Applicable
Fertilization Broadcast Spawning
Birth Type Broadcast_spawning

Across Clupeidae, mating is typically group-based broadcast spawning: many males and females release gametes into the water in schools or spawning aggregations, with no pair bonds or parental care. Timing and spawning locations vary widely among species and regions.

Behavior & Ecology

Social School Group: 10000
Activity Diurnal, Crepuscular, Cathemeral
Diet Filter Feeder Copepods (dominant zooplankton prey for many Clupeidae; often a key driver of growth and schooling distribution)
Seasonal Migratory 1,243 mi

Temperament

Strongly gregarious; cohesion increases under predation pressure
Generally non-territorial and low aggression within schools
Risk-averse; rapid collective escape responses and frequent school re-shaping
Often migratory or seasonally aggregative; extent varies widely among species and habitats
Foraging ranges from continuous plankton filtering to more selective particulate feeding, depending on species and prey

Communication

none documented as primary communication; sounds mainly incidental during movement
lateral-line detection of water movement for alignment and spacing
visual cues (silvery flashes, neighbor orientation) for synchronized turning
hydrodynamic signaling via rapid speed/heading changes propagating through the school
chemical cues for conspecific recognition and predator/alarm responses in some species
school geometry adjustments (tight balling, milling, layering) as group-level signaling

Habitat

Biomes:
Marine Freshwater Wetland Tropical Rainforest Tropical Dry Forest Mediterranean Temperate Forest Boreal Forest (Taiga) Tundra Desert Hot Desert Cold +5
Terrain:
Coastal Island Riverine
Elevation: -13780 in

Ecological Role

Mid-trophic pelagic planktivore and foundational forage-fish group (major conduit of plankton production to higher predators and human fisheries)

Transfers energy from phyto/zooplankton to predators (seabirds, marine mammals, larger fishes) Stabilizes and shapes plankton communities via intense grazing (top-down control on plankton blooms in some systems) Supports major commercial fisheries and food webs (high-biomass schooling prey base) Nutrient cycling via excretion and mass schooling (local fertilization and redistribution of nitrogen/phosphorus) Carbon transport/export through feeding and fecal pellet production (can enhance carbon sinking from surface waters)

Diet Details

Main Prey:
Zooplankton Krill Cladoceran Amphipods Crustaceans Fish eggs and fish larvae Zooplankton +1
Other Foods:
Phytoplankton Microalgae and suspended primary producers

Human Interaction

Domestication Status

Wild

Clupeidae (herrings, sardines, shads, menhadens, sprats) are not domesticated. People mainly catch wild fish and process them fresh, salted, smoked, canned, or for fishmeal/oil. Small-scale farming, holding for research, bait supply, and short-term live storage happen, but no long-term breeding like in carp or salmon. Hatchery work for some shads aids conservation, not domestication.

Danger Level

Low
  • Foodborne illness risks if mishandled: histamine (scombroid-like) poisoning can occur in clupeids when temperature control fails; also general bacterial contamination risks
  • Parasites can be present in some species/areas (risk reduced by proper cooking/freezing practices)
  • Biotoxins/contaminants may accumulate depending on local blooms/pollution (e.g., algal toxins), varying strongly by region and season
  • Choking hazard from fine bones, especially in whole/small fish preparations
  • Allergic reactions in sensitive individuals (fish allergy)
  • Minor injuries to fishers/handlers (cuts from gill covers/spines, hooks, slips on decks)

As a Pet

Not Suitable as Pet

Legality: Sardines are usually legal to keep where local rules allow native fish, but rules often limit taking wild fish, moving live bait, invasive species laws, and animal welfare. Some species are protected or have seasonal limits; laws vary by place.

Care Level: Expert Only

Purchase Cost: Up to $25
Lifetime Cost: $500 - $20,000

Economic Value

Uses:
Industrial capture fisheries Small-scale/coastal fisheries Food processing (canning/smoking/salting) Bait fisheries Fishmeal and fish oil production Aquaculture feed inputs Ecosystem services (forage base supporting higher-value fisheries)
Products:
  • canned sardines/herrings
  • fresh/chilled whole fish
  • frozen products
  • salted and smoked fish
  • roe products (in some regional markets)
  • fishmeal
  • fish oil (including omega-3 rich oils)
  • bait (fresh/frozen/live where permitted)
  • fertilizer/soil amendments (from byproducts in some regions)

Relationships

Predators 8

Tuna
Tuna Thunnus spp.
Mackerel Scomber spp.
Seabirds Morus spp.; Phalacrocorax spp.; Fratercula spp.
Dolphin
Dolphin Delphinidae
Seals and sea lions Pinnipedia
Cod
Cod Gadus spp.
Salmon
Salmon Oncorhynchus spp.; Salmo spp.
Shark
Shark Carcharhinus spp.

Related Species 12

Atlantic herring Clupea harengus Shared Family
Pacific herring Clupea pallasii Shared Family
European pilchard Sardina pilchardus Shared Family
South American pilchard Sardinops sagax Shared Family
Japanese sardine Sardinops melanostictus Shared Family
Round sardinella Sardinella aurita Shared Family
European sprat Sprattus sprattus Shared Family
American shad Alosa sapidissima Shared Family
Alewife
Alewife Alosa pseudoharengus Shared Family
Atlantic menhaden Brevoortia tyrannus Shared Family
Pacific menhaden Ethmidium maculatum Shared Family
Scaled sardine Harengula jaguana Shared Family

Ecological Equivalents 5

Animals that fill a similar ecological role in their ecosystem

Anchovies
Anchovies Engraulidae Occupy a similar niche to small, schooling pelagic/neritic planktivores; often co-occur with clupeids and are harvested together as forage fish.
Capelin Mallotus villosus Cold-water, schooling planktivore that forms massive spawning aggregations and serves as key prey for seabirds and larger fish; functionally similar to many clupeids in food webs.
Sand lance Ammodytidae Schooling forage fishes in coastal upwelling and temperate systems; overlap as a prey base for seabirds, marine mammals, and predatory fishes, even though feeding mode and habitat use differ.
Round herring Etrumeus teres Sardine-like round herring. Small pelagic, schooling planktivore commonly confused with sardines in fisheries and markets; occupies a similar trophic role in many regions.
Small mackerels Scomber spp. Juvenile mackerels overlap strongly in size and schooling behavior with clupeids in many ecosystems, consume similar zooplankton, and also prey on small clupeids.

Types of Sardines

23

Explore 23 recognized types of sardines

European sardine (European pilchard) Sardina pilchardus
South American pilchard / sardine Sardinops sagax
Japanese sardine Sardinops melanostictus
Round sardinella Sardinella aurita
Flat sardinella Sardinella maderensis
Indian oil sardine Sardinella longiceps
European sprat Sprattus sprattus
Brisbane/blue sprat Spratelloides gracilis
Atlantic herring Clupea harengus
Pacific herring Clupea pallasii
American shad Alosa sapidissima
Allis shad Alosa alosa
Twaite shad Alosa fallax
Alewife (river herring) Alosa pseudoharengus
Blueback herring (river herring) Alosa aestivalis
Atlantic menhaden Brevoortia tyrannus
Gulf menhaden Brevoortia patronus
Scaled sardine Harengula jaguana
Thread herring Opisthonema oglinum
Pacific thread herring Opisthonema libertate
Gizzard shad Dorosoma cepedianum
Pacific sardine Sardinops caeruleus

Quick Take

Sardines are a group of herring fish that encompasses five distinct genera and at least 21 unique species. One of the most common sardines used in cooking and food preparation is the Pacific Sardine, also known as Sardinops sagax.

Sardines are small fish that typically do not exceed 12 inches in length and can weigh as little as three ounces each. However, some larger herring varieties, such as the American Shad, can reach as much as 12 pounds in weight.

European fisheries refer to these fish as pilchards. In fact, the UK and other European nations define them as juvenile pilchards that are less than 6 inches (15 cm) in length. Other varieties include the South American Pilchard, the Bali Sardinella, and the Indian Oil Sardine.

These fish all have silvery scales and one small dorsal fin. They swim in large schools that can stretch for miles. They are found in almost every area of the world, including the Atlantic and Pacific Oceans, the Mediterranean Sea, and the waters around Morocco and other nations.

Detailed infographic about sardines showing their anatomy, food chain role as a prey species for dolphins and whales, and the commercial fishing impact comparing current biomass to sustainable levels.
They fuel the ocean's apex predators and protect your heart health, but a plummeting biomass threatens the entire marine food chain. Discover why these tiny powerhouses are more than just a pantry staple. © A-Z Animals

Sardines vs. Anchovies?

School of Swimming Anchovies

Anchovies are pretty similar to sardines, although they have a reddish color when prepared.

Sardines and Anchovies may seem alike in many ways. Both are smaller fish that can be canned for consumption. They are both used heavily in cooking as a source of omega-3 fatty acids, vitamins, and minerals.

However, Sardines have a prominent lower jaw and white flesh when cooked or salted. In contrast, Anchovies have smaller mouths and are a reddish color when smoked, salted, or otherwise prepared.

5 Sardines Facts

  • Named for the island of Sardinia in the Mediterranean, where these fish were plentiful.
  • More than 21 unique species of sardines are part of the herring family.
  • Found all over the world, from the North Atlantic and Pacific Oceans to the Mediterranean Sea, the Indian Ocean, and the waters around South America.
  • Normally canned as food in olive oil, water, tomato juice, or other liquids.
  • Excellent source of omega-3 fatty acids, which promote heart health.

Evolution

Diver and large school of sardines

The earliest ancestors of sardines first emerged during the Late Jurassic period.

Sardines are Clupeiformes, a classification of fish that can be traced back to the Late Jurassic, a period that occurred between 145 and 200 million years ago. The group includes anchovies, which belong to the Engraulidae family, as well as the Clupeidae family. The latter group consists of herrings, sardines, and shads and dates back to the Paleogene era, i.e., between 23 and 66 million years ago.

Members of the Clupeidae family are noted for absent or reduced teeth, gleaming scales, the presence of a single dorsal fin in the majority of cases, and a love of plankton. They are also impressively prolific producers and occupy a rather low position on the food chain.

Classification and Scientific Name

Sardines is a broadly defined name for several small herring fish. There are at least 21 different species across five different genera.

Species

There are over 21 different species of sardine fish throughout the world. Some of the most well-known are:

  • The European Pilchard, or Sardina pilchardus: Occasionally capable of growing to 11 inches, the European Pilchard can be found in the Atlantic, Black Sea, and the Mediterranean.
  • The Freshwater Sardine (Sardinella tawilis): Endemic to the Philippines, this fish, which can be recognized by its triangular dorsal fin, is capable of growing to 6 inches. It is also known as the freshwater herring.
  • The Rainbow Sardine (Dussumieria acuta): Mostly found in Southeast Asia, this fish, also known as the rainbow herring, is capable of growing to 8 inches.
  • The South American Pilchard (Sardinops sagax): This species can grow up to 16 inches and can be found in the Indo-Pacific and the eastern Pacific.
  • The White Sardine (Escualosa thoracata): Generally found in Thailand, Indonesia, and extending to Australia, this species is capable of growing to a maximum length of 4 inches.

Appearance

Quite a few herring fish are referred to as sardines when they are juveniles. These fish are typically less than 12 inches long and have an elongated body with silvery scales. They typically have no scales on their heads and only one small dorsal fin on the top of their body.

Because more than 20 fish species are considered sardines, this fish’s weight can vary substantially. Some types weigh as little as two or three ounces, while the largest sardines may reach up to one pound.

Bronze whaler shark swimming through a large sardine bait ball, looking to feed during the sardine run, east coast of South Africa.

Distribution, Population, and Habitat

The numerous species are found in saltwater oceans, seas, and other bodies of water throughout the world. Schools of these fish frequent the Atlantic Ocean and the waters around California and Mexico. They are also common around South America, Europe, Africa, and Asia.

The exact numbers of each species are unknown. However, populations are measured in metric tons to determine whether these creatures are being overfished. For example, the Pacific Sardine’s population is currently estimated at more than 30,158 metric tons in biomass. While that may sound like a lot, the threshold required to sustain commercial fishing is 150,000 metric tons. As a result, the NOAA currently classifies the Pacific Sardine as overfished.

Predators

Animal Facts: Penguins

Penguins enjoy sardines, which they consider to be a nourishing snack.

Many marine animals and birds rely on these fish as a key component of their diet. Dolphins, seals, and Blue Whales are examples of ocean mammals that eat these fish. Penguins, terns, Brown Pelicans, and other seafaring birds also love to scoop herrings out of the water for a snack.

Humans are another significant predator of sardines. Because these fish only swim between 0 and 200 meters below the surface, our impact on their habitats is relatively minor compared to other sea life. However, they are highly sought after in Europe, Asia, and India as a source of nutrition. Overfishing and commercialization are major risks to the worldwide sardine population.

Diet

Sardines typically eat zooplankton and phytoplankton, which are microscopic organisms found in the Earth’s oceans. Plankton can include the small eggs of larger fish species as well as tiny crustaceans and other creatures that cannot swim along the ocean currents.

Reproduction and Lifespan

These fish can spawn as early as one to two years of age. Females can have between 10,000 and 40,000 eggs available for fertilization. Initially, scientists believed that sardines spawned only once or twice per season, but recent studies indicate that it may be much more frequent.

Fishing and Cooking

Sardines on white background

Sardines are widely eaten all around the globe.

Sardines are a common source of nutrition around the world. In fact, global commercial fishing for sardines has varied widely, but in recent years annual catches have often fallen below three million metric tons due to population declines and fishery closures.

Sardines are frequently canned in olive oil, which helps the meat remain moist and tender and preserves its rich flavor. Cheaper sardines can be kept in tin cans filled with water, tomato juice, or other liquids. However, these approaches can leave the meat soggy or slimy.

The fish can also be smoked or salted to preserve their dense white flesh. Many people choose to eat them straight out of the can on crackers with tomatoes or onions.

Population

Marlins hunting on sardines or makerels in Baja California Sur

Pacific sardines are believed to have a population that is equivalent in weight to 27 metric tons.

Estimating the worldwide population of sardines is challenging because of the diverse regions where they live. The large number of species included in the pilchard or herring family also makes a numerical count difficult. Instead, sardine populations are often measured in metric tons.

For example, Pacific sardines, which are one of the most popular options for canning, salting, and cooking, have an estimated population equivalent to more than 27 metric tons in weight.

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Sources

  1. Wikipedia / Accessed December 24, 2020
  2. Animal Diversity Web / Accessed December 24, 2020
  3. Food & wine / Accessed December 24, 2020
  4. Health Essentials / Accessed December 24, 2020
  5. Fishbase / Accessed December 24, 2020
  6. Tin Can fish / Accessed December 24, 2020
Dana Mayor

About the Author

Dana Mayor

I love good books and the occasional cartoon. I am also endlessly intrigued with the beauty of nature and find hummingbirds, puppies, and marine wildlife to be the most magical creatures of all.
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Sardines FAQs (Frequently Asked Questions)

Sardines, also called pilchards in Europe, are one of several fish that are part of the herring family. In fact, five different genera include more than 16 distinct species of sardines. As a result, these vertebrates are found all over the world, from the North Atlantic to South America, Asia, and India.

They are generally small, but lengthy fish that measure between six and 12 inches long. However, some species, such as the South American pilchard, can reach more than 15 inches long. They usually have one dorsal fin and a silvery color with blue eyes.