T
Species Profile

Tussock Moth Caterpillar

Erebidae

Hairy larvae, big forest impact
Satyashutter/Shutterstock.com

Tussock Moth Caterpillar Distribution

Click a location to explore more animals from that region

Invasive Species
Loading map...
Macro of a white-marked tussock moth caterpillar. The caterpillar is facing the camera, center frame, on a leaf that has holes in it. The caterpillar is incredible! it has large false eyes that make it appear to be animatronic, It is really otherworldly looking. The top of its back is brown. White hairs (setae) extend from the side of its body, which is white.

At a Glance

Family Overview This page covers the Tussock Moth Caterpillar family as a group. Stats below are general traits shared across the family.
Also Known As tussock caterpillar, tussock moth larva, hairy caterpillar, bristly caterpillar
Diet Herbivore
Activity Nocturnal+
Lifespan 365 years
Weight 0.01 lbs
Status Not Evaluated
Did You Know?

Erebidae is huge: adults range roughly 10-180 mm wingspan, showing extreme size diversity within one moth family.

Scientific Classification

Family Overview "Tussock Moth Caterpillar" is not a single species but represents an entire family containing multiple species.

“Tussock moth caterpillar” refers to the larvae of tussock moths (subfamily Lymantriinae). They are typically hairy, often brightly patterned caterpillars that feed on leaves of trees and shrubs. Some species are important defoliators and many have irritating hairs that can cause rashes.

Kingdom
Animalia
Phylum
Arthropoda
Class
Insecta
Order
Lepidoptera
Family
Erebidae

Distinguishing Features

  • Dense hairs with tufts (“tussocks”)
  • Hair pencils projecting near head or tail
  • Often causes skin irritation on contact
  • Primarily leaf-feeding on trees and shrubs

Physical Measurements

Males and females differ in size

Length
1 in (0 in – 3 in)
Weight
0 lbs (0 lbs – 0 lbs)
0 lbs (0 lbs – 0 lbs)
Poisonous

Appearance

Primary Colors
Secondary Colors
Skin Type Hairy setae
Distinctive Features
  • Larval length ranges roughly 0.8-7 cm across the group.
  • Dense defensive setae form "tussocks" on dorsal segments.
  • Hair pencils or tufts often project at head and tail.
  • Coloration highly variable: cryptic browns to vivid warning patterns.
  • Hairs can irritate skin; reactions vary among species.
  • Feeding mostly on tree and shrub leaves; host ranges vary.
  • Some species are major defoliators during outbreak years.
  • Early instars may feed gregariously; later instars often solitary.
  • Many rest openly on foliage; others hide by day.
  • Overwintering stage varies: eggs, larvae, or pupae depending species.
  • Setae may detach easily, aiding defense against predators.
  • Body surface largely hidden by hairs; true skin rarely visible.

Sexual Dimorphism

Sexual dimorphism is mainly expressed in adults, not larvae: males are often smaller with strongly feathered antennae, while females can be larger and sometimes reduced-wing or flightless in some lineages.

  • Often smaller-bodied adult moths, more active fliers.
  • Strongly bipectinate antennae for detecting female pheromones.
  • Duller wing coloration is common, but varies by species.
  • Often larger-bodied adults with heavier abdomens for egg production.
  • Some species have reduced wings or flightlessness.
  • Antennae usually less feathery than males.

Did You Know?

Erebidae is huge: adults range roughly 10-180 mm wingspan, showing extreme size diversity within one moth family.

Across Erebidae, caterpillars span about 5-100 mm long; tussock moth (Lymantriinae) larvae commonly sit mid-range.

Many tussock moth caterpillars carry barbed "urticating" hairs that can cause rashes, eye irritation, or respiratory discomfort.

Some lymantriine larvae are serious forest and orchard defoliators, especially during outbreak years when populations boom.

Life cycles vary widely: from a few weeks in warm climates to 1-3+ years with overwintering diapause.

Early instars of many tussock moths feed in groups, while older larvae often spread out to reduce competition.

Erebidae also includes tiger moths and underwings; chemical defenses and warning colors evolved repeatedly across the family.

Unique Adaptations

  • Detachable, barbed setae form a physical defense that can lodge in skin and mucous membranes.
  • Bold stripes, tufts, and contrasting "tussocks" function as warning signals, often paired with chemical deterrents.
  • Some Erebidae sequester plant toxins or produce defensive chemicals; aposematic larvae advertise unprofitability.
  • Silk and hooks on abdominal prolegs help caterpillars grip foliage and move efficiently through tree canopies.
  • Diapause flexibility (egg, larva, or pupa depending on species) lets populations survive cold, drought, or poor food years.

Interesting Behaviors

  • Young larvae of many tussock moths feed gregariously, skeletonizing leaves before dispersing as they grow.
  • Caterpillars often "balloon" on silk threads, letting wind disperse them to new host plants.
  • When disturbed, many species arch, thrash, or expose brighter patches, emphasizing their irritating hairs.
  • Host use ranges from broad generalists to strict specialists tied to particular tree or shrub lineages.
  • Population dynamics can swing dramatically; outbreaks may cause widespread defoliation, then crash from disease and predators.

Cultural Significance

Tussock moth caterpillars shape forestry and urban tree care through periodic defoliation, while their irritating hairs drive public-health guidance. Erebidae also fuels folklore-especially "winter" caterpillar weather sayings-and inspires pest-control history and quarantine policy.

Myths & Legends

In North American folklore, the "woolly bear" caterpillar's banding is said to foretell how harsh the coming winter will be.

Sufi and South Asian poetic tradition uses the moth and candle as a legend-like moral image of love, devotion, and self-sacrifice.

A widely repeated historical tale recounts the accidental North American release of the spongy moth during silk experiments, becoming a cautionary story about introduced pests.

You might be looking for:

White-marked Tussock Moth Caterpillar

26%

Orgyia leucostigma

Common North American tussock moth larva with distinctive toothbrush-like tufts and irritant hairs; often found on hardwood trees in towns and forests.

Rusty Tussock Moth Caterpillar (Vapourer)

22%

Orgyia antiqua

Widespread Palearctic species; caterpillar has prominent hair pencils and tufts; feeds on many shrubs and trees, frequently seen in gardens.

Douglas-fir Tussock Moth Caterpillar

18%

Orgyia pseudotsugata

Forest pest in western North America; larvae defoliate Douglas-fir and true firs during outbreaks; hairs can cause skin irritation.

Spongy Moth Caterpillar (formerly “Gypsy moth”)

17%

Lymantria dispar

Notorious defoliator; caterpillars have paired blue and red dorsal spots; a tussock-moth relative often implicated when people mention hairy tree caterpillars.

View Profile

Life Cycle

Birth 300 larvas
Lifespan 365 years

Lifespan

In the Wild
25–1095 years
In Captivity
25–56 years

Reproduction

Mating System Promiscuity
Social Structure Solitary
Breeding Season Spring-summer; year-round in tropics; variable
Breeding Pattern Transient
Fertilization Internal Fertilization
Birth Type Internal_fertilization

Across Erebidae, adults are typically solitary; females attract mates with pheromones and pair briefly. Males often mate multiple times, while female remating ranges from single to multiple depending on species. No lasting bonds; females oviposit without helpers.

Behavior & Ecology

Social Aggregation Group: 5
Activity Nocturnal, Diurnal, Crepuscular
Diet Herbivore tree leaves
Seasonal Hibernates

Temperament

Cryptic
Defensive
Startle-prone
Aposematic
Gregarious

Communication

ultrasonic clicks
wing rustle sounds
sex pheromones
aggregation pheromones
silk trail cues
substrate vibrations
tactile contact
visual startle displays
chemical defenses

Habitat

Biomes:
Tropical Rainforest Tropical Dry Forest Savanna Desert Hot Desert Cold Mediterranean Temperate Grassland Temperate Forest Temperate Rainforest Boreal Forest (Taiga) Tundra Alpine Freshwater Wetland +8
Terrain:
Mountainous Hilly Plateau Plains Valley Coastal Island Riverine Volcanic Karst Rocky Sandy Muddy +7
Elevation: Up to 14763 ft 9 in

Ecological Role

Primary herbivores, occasional detritivores; key prey and periodic defoliators

food web support vegetation regulation nutrient cycling forest succession effects energy transfer to predators

Diet Details

Other Foods:
Tree leaves Shrub leaves Herbaceous plants Grasses Lichen Algae films Crop foliage +1

Human Interaction

Domestication Status

Wild

Erebidae (including tussock moths, Lymantriinae) have not been domesticated. Humans primarily interact with them as wild insects-often as forestry/agricultural pests, occasional laboratory study organisms, and education/display specimens rather than for sustained husbandry.

Danger Level

Moderate
  • urticating hairs causing dermatitis
  • eye irritation from airborne setae
  • allergic reactions in sensitized people
  • nuisance outbreaks near homes
  • tree and shrub defoliation impacts

As a Pet

Not Suitable as Pet

Legality: Often unregulated; pest species may be quarantined/restricted.

Care Level: Experienced

Purchase Cost: Up to $30
Lifetime Cost: $5 - $150

Economic Value

Uses:
Forestry Agriculture Research Education
Products:
  • specimens

Relationships

Predators 8

Black-capped chickadee
Black-capped chickadee Poecile atricapillus
Common cuckoo Cuculus canorus
Yellow warbler Setophaga petechia
Tachinid flies Tachinidae
Ichneumon wasp Ichneumonidae
Compsilura tachinid fly Compsilura concinnata
Ground beetle
Ground beetle Carabidae
Orb-weaver spider
Orb-weaver spider Araneidae

Related Species 8

White-marked tussock moth Orgyia leucostigma Shared Family
Vapourer Orgyia antiqua Shared Family
Douglas-fir tussock moth Orgyia pseudotsugata Shared Family
Spongy moth
Spongy moth Lymantria dispar Shared Family
Brown-tail moth Euproctis chrysorrhoea Shared Family
Fall webworm moth Hyphantria cunea Shared Family
Owlet moths Noctuidae Shared Family
Giant silk moths
Giant silk moths Saturniidae Shared Family

Ecological Equivalents 5

Animals that fill a similar ecological role in their ecosystem

Eastern tent caterpillar Malacosoma americanum Tree-defoliating larva. Outbreaks can strip foliage from hardwoods.
Oak processionary moth caterpillar Thaumetopoea processionea Gregarious tree-feeding defoliator with urticating hairs that cause dermatitis.
Puss caterpillar
Puss caterpillar Megalopyge opercularis Hairy larva with irritating (stinging) defenses; feeds on many tree species.
Pine sawfly larvae Diprionidae Conifer defoliators. Larval swarms can cause major needle loss.
Io moth caterpillar Automeris io Stinging spines deter predators. Larva feeds on broadleaf shrubs and trees.

Types of Tussock Moth Caterpillar

11

Explore 11 recognized types of tussock moth caterpillar

White-marked tussock moth Orgyia leucostigma
Vapourer (rusty tussock moth) Orgyia antiqua
Douglas-fir tussock moth Orgyia pseudotsugata
Spongy moth (formerly gypsy moth) Lymantria dispar
Pale tussock Dasychira pudibunda
Brown-tail moth Euproctis chrysorrhoea
Satin moth Leucoma salicis
Fall webworm moth Hyphantria cunea
Yellow woolly bear moth Spilosoma virginica
Black witch moth
Black witch moth Ascalapha odorata
Fruit-piercing moth
Fruit-piercing moth Eudocima fullonia

The tussock moth caterpillar is the larval stage of a tussock moth, which is a type of moth in the family Erebidae. Tussock moth caterpillars are typically fuzzy and have tufts of hair, or tussocks, on their bodies. They can be found in a variety of colors, including white, yellow, orange, and brown. They can be found feeding on the leaves of trees and shrubs, and are considered pests by some farmers and gardeners. This is due to their ability to defoliate trees, as well as entire forests! Keep reading to learn more about these hungry caterpillars!

Five Furry Facts about Tussock Moth Caterpillars

  • Tussock moth caterpillars have tufts of hair or tussocks along their bodies, which is their most distinctive feature.
  • They are herbivorous, and they feed on the leaves of various trees and plants.
  • Vapourer moths (Orgyia antiqua) do not spin cocoons, metamorphosing directly into adult moths.
  • They are active during the day and they tend to rest at night.
  • They are preyed upon by a variety of predators, including birds, mammals, insects, and some parasitic wasps.
Macro of a hickory tussock caterpillar. It is mostly gray with mostly white hairs stuck out from its body, giving it a fuzzy appearance. There are two distinct black tufts, on toward the from, and one toward the back. Against medium brown background

Tussock moth caterpillars are hairy, like this hickory tussock moth caterpillar (Lophocampa caryae).

Scientific Name

Tussock moth caterpillars share their scientific names with their adult moth forms. Tussock moths comprise a large group of moths, of which there are many different species. However, they all belong to the same family, Erebidae. Each species has its own scientific name. The spongy moth (Lymantria dispar), the Douglas-fir tussock moth (Orgyia pseudotsugata), and the White-marked tussock moth (Orgyia leucostigma) are all in the family Erebidae. Erebidae is Greek and loosely translates to out of the darkness. This seems like a fitting name for adult moths, who tend to fly at night. There are 350 genera of tussock moths comprised of more than 2700 separate species! New species continue to be discovered and classified.

Macro of a white-marked tussock moth caterpillar. The caterpillar is facing the camera, center frame, on a leaf that has holes in it. The caterpillar is incredible! it has large false eyes that make it appear to be animatronic, It is really otherworldly looking. The top of its back is brown. White hairs (setae) extend from the side of its body, which is white.

The white-marked tussock moth (Orgyia leucostigma) is one of over 2000 tussock moth species.

Tussock Moth Caterpillars: Appearance

Tussock moth caterpillars are small, fuzzy insects that can vary in color from yellow to brown to black. Many tussock moth caterpillars have urticating hairs, which cause an unpleasant reaction in some individuals. However, reactions tend to be mild, and are the result of an allergic reaction. Tussock moth caterpillars are not venomous. They have tufts of hair, called setae (seta, singularly) or tussocks along their bodies, which are their most distinctive feature. Some species have distinctive markings or patterns on their bodies as well.

They generally range from 0.5 to 2 inches in length. Tussock moth caterpillars are active during the day, tending to rest at night. Some species spin cocoons and metamorphose into adult moths, while others do not form cocoons. Species of the genus Orgyia, such as the vapourer moth (Orgyia antiqua) do not spin cocoons. Instead, they form a pupal case around their body in which they metamorphose into adult moths. Known as a naked pupae. It is a characteristic feature of the genus Orgyia.

Macro of a vapourer moth caterpillar. The caterpillar is primarily dark brown covered in red orange and yellow spots from which hairs protrude. From the midsection of its back on four distinct, separate tufts of hair are visible. These tufts are cream to light yellow in color. The caterpillar also has distinct feathery antennae. It is on a green stem, horizontal across the frame with its face facing right.

Species of the genus Orgyia, such as the vapourer moth (Orgyia antiqua) do not spin cocoons. Instead, they form a pupal case around their body.

Tussock Moth Caterpillars: Habitat

Tussock moths are found in a wide variety of habitats, including forests, woodlands, meadows, and gardens. They are found throughout much of Africa, Asia, Europe, and North America. Some species, such as the spongy moth (Lymantria dispar), are known to occur in both urban and rural areas. They are also known to occupy a wide range of elevations, from sea level to alpine habitats. The caterpillars of some tussock moth species can be found on a variety of trees and plants, including oak, willow, apple, cherry, and many others. Adult moths are generally nocturnal and are attracted to lights
Tussock moth caterpillars are generally diurnal, actively feeding throughout the day. They feed on the leaves of various trees and plants during the day and rest at night. Some species, such as the gypsy moth (Lymantria dispar) and the vapourer moth (Orgyia antiqua) are known to migrate in large groups, which can make their effects on plants even more severe. However, it’s important to note that there could be some variations between different species of tussock moths, as some species might be more active at night or crepuscular.

Tussock Moth Caterpillars: Diet

Tussock moth caterpillars are herbivorous and they feed on the leaves of various trees and plants. They have a wide range of host plants and they can be found on deciduous and coniferous trees, shrubs, and other plants. The host plants vary depending on the species of tussock moth. For example, the caterpillars of the spongy moth (Lymantria dispar) favor oak, willow, apple, cherry, and many other deciduous trees. The caterpillars of the vapourer moth (Orgyia antiqua) feed on deciduous trees such as oak, birch, and willow, and also on conifers like spruce, pine, larch, and Douglas fir.

Some tussock moth caterpillars are generalist feeders that will eat what is available. Of note, many species of tussock moth caterpillars are considered pests. This is thanks to their ability to defoliate entire forests! The spongy moth (Lymantria dispar) and the Douglas-fir tussock moth (Orgyia pseudotsugata) are two species that can completely decimate ecosystems.

A gray and brown spongy moth caterpillar with hair sticking out from every direction. In the photograph the caterpillar appears to be facing the left with two false black eyes visible toward the left part of the frame along the caterpillar’s back are four pairs of black dots sprouting black hairs and then six pairs of red dots sprouting black hairs. The caterpillar skin is mottled from light tan to dark brown. The caterpillar is perched on a green leaf with yellow veins.

Spongy moth caterpillars (Lymantria dispar) favor oak, willow, apple, cherry, and many other deciduous trees.

Tussock Moth Caterpillars: Control Measures

In attempting to avoid the destruction of forests and other habitats where tussock moth caterpillars are found, there are a variety of methods that are being used to control or eradicate tussock moths. Some of these include:

  • Biological control: This involves the use of natural predators or pathogens to reduce the population of tussock moths. For example, parasitic wasps are introduced to target the eggs or larvae of the moths.
  • Chemical control: Insecticides can be used to kill tussock moths. However, this method can be harmful to other non-target organisms and must be used with care.
  • Cultural control: Though it seems counterintuitive, this method involves removing or destroying the host plants of tussock moths, as well as maintaining healthy trees and plants that can resist infestation.
  • Trapping: Some farmers use pheromone traps to capture the moths, which is a more targeted approach to control.
  • Removal: This includes removing the caterpillars by hand or using a vacuum to suck them up.
Macro of a spongy moth caterpillar on an almost completely destroyed oak leaf. The caterpillar has presumably eatenmozt of the leaf. What's left of the leaf is green. The caterpillar is brownish/gray with stiff black hairs sticking out from its body. On the top of its back is sports four pairs of blue spots and six pairs of red.

Spongy moth caterpillars, like the one in the photo, can wreak havoc on ecosystems.

It’s important to note that the most effective control method depends on the specific species of tussock moth and their specific location and environment. A combination of methods is often necessary to achieve the desired results.

Tussock Moth Caterpillars: Predators

Tussock moth caterpillars face a variety of predators, including birds, mammals, and insects. Birds such as chickadees, nuthatches, and woodpeckers are known to feed on tussock moth caterpillars. Many species of mammals, including squirrels, chipmunks, and deer, also feed on tussock moth caterpillars. Additionally, insects, such as lady beetles, lacewings, and wasps, prey on tussock moth caterpillars. Parasitic Braconid wasps, like the Cotesia melanoscela and Apanteles melanoscelus, lay their eggs on the tussock moth caterpillars, and when the wasp larvae hatch, they feed on the inside of the caterpillars, eventually killing them. Some species, like the white-marked tussock moth (Orgyia leucostigma), have a symbiotic relationship with ants. The ants provide a distraction allowing the caterpillars to escape detection, generally from bigger, meaner ants. In exchange, the protector ants receive the sugars and amino acids secreted by the caterpillar’s dorsal nectary organ, a mutualistic relationship known as myrmecophily.

Tussock Moth Caterpillars: Threats

Tussock moth caterpillars face a variety of threats, including predation, disease, and exposure to extreme environmental conditions. They are also affected by other naturally occurring phenomena such as parasitism, viral and fungal diseases, and competition for food. Additionally, tussock moth caterpillars can be affected by human activities such as logging, urbanization, and pesticide use. Logging and urbanization can cause the destruction of their natural habitats, leading to a decline in their population. Pesticides can also be harmful to tussock moth caterpillars, both directly and indirectly, by killing them or by reducing the population of their prey or host plants. Climate change is also a threat to many species of tussock moths, as it can change the timing of their life cycle, and make them vulnerable to new pathogens, predators, and parasites.

Conservation Status

The conservation status of tussock moth caterpillars varies depending on the species. Some species are common and widespread, while others are considered rare or endangered. The International Union for Conservation of Nature (IUCN) Red List of Threatened Species is a widely recognized system for assessing the conservation status of species. However, tussock moth caterpillars are not specifically listed in the IUCN Red List, making it difficult to give an overall status for them as a group.

Some species of tussock moths such as the Gypsy Moth are considered pests and are targeted for control by various methods. On the other hand, other species of tussock moths are not considered pests. Some others are even considered indicator species, particularly those in the subfamily Lymantriinae, used for determining the health of certain habitats. For accurate current information regarding specific tussock moth species, best practices suggest consulting a local university, agricultural extension center, or conservation group.

Macro if milkweed tussock caterpillar on a bright green leaf. Thecterpillars is quite fuzzy. It has whitish/gray hair protruding from the bottom part of its body. The middle is black, and the top is striped mpstlyorng with some gray and black. Looks like a parade float.

For current and accurate information regarding specific tussock moth species, like this milkweed tussock, consult a local university, agricultural extension center, or conservation group.

Lifecycle

The lifecycle of tussock moth caterpillars typically begins with the adult moths laying eggs on the leaves of trees and plants. The eggs are usually laid in clusters, hatching within a few weeks. Once they hatch, the caterpillars begin feeding on the leaves of their host plants. As they grow, they will molt several times, shedding their exoskeleton to reveal a new, larger one. The shedding of their exoskeleton is called an instar. Caterpillars will typically molt 5-6 times, or have 5-6 instars.

Tussock moth caterpillars are active during the day and they tend to rest at night. They continue to feed and grow until they reach their full size, which can take anywhere from a few weeks to several months, depending on the species. Once they reach maturity, the caterpillars will spin a cocoon or form a pupal case around their body in which they will metamorphose into adult moths. The adult moths emerge from the cocoon or pupal case after a period of time, which can be a few days to several weeks, depending on the species. In their adult form, they mate and lay eggs, beginning the cycle anew.

Macro of Vapourer Moth (Orgyia antiqua) eggs on peanut leaf. There are 100s of eggs in the frame. The eggs look like c=the cereal Cheerios, but with indistinct center holes.

Tussock caterpillars begin life in eggs, like these vapourer moth (Orgyia antiqua) eggs.

Lifespan

The lifespan of tussock moth caterpillars can vary depending on the species and environmental conditions. The entire life cycle, from egg to adult, can take from a few weeks to several months. The eggs hatch within a few weeks, and the caterpillars typically go through several stages of growth, shedding their exoskeleton multiple times before reaching maturity. The mature caterpillars will then spin a cocoon or form a pupal case and metamorphose into adult moths. Adult moths live for a relatively short period of time, usually a few days to a few weeks.

Population

Population estimates for tussock moth caterpillars can vary depending on the species and the location. Some species are considered to be common and widespread, while others are considered to be rare or endangered. However, it’s hard to give a general estimate of population size for tussock moth caterpillars as it can fluctuate greatly depending on factors such as the availability of host plants, weather conditions, and predation. Additionally, tussock moth populations can fluctuate greatly from year to year and can be impacted by various factors such as weather, predators, and disease. Also, tussock moths are not specifically monitored by any agency, so there is no specific data on population estimates

Up Next:

View all 608 animals that start with T

Sources

  1. researchgate.net / Accessed January 25, 2023
  2. ufl.org / Accessed January 25, 2023
  3. insectidentification.org / Accessed January 25, 2023
  4. caterpillaridentification.org / Accessed January 25, 2023
  5. wikipedia.org / Accessed January 25, 2023
  6. amentsoc.org / Accessed January 25, 2023
  7. wpmucdn.com / Accessed January 25, 2023
Kathryn Koehler

About the Author

Kathryn Koehler

Kathryn Koehler is a writer at A-Z-Animals where her focus is on unusual animals, places, and events. Kat has over 20 years of experience as a professional writer and educator. She holds a master's degree from Vanderbilt University. When she is not writing for A-Z-Animals, Kat enjoys puttering in her garden, baking deliciously healthful treats for her family, and playing with her two rescue mutts, Popcorn and Scooter. She resides in Tennessee.

Thank you for reading! Have some feedback for us?


Tussock Moth Caterpillar FAQs (Frequently Asked Questions)

No! Tussock moth caterpillars are not venomous.  Many tussock moth caterpillars have urticating hairs which can cause an unpleasant reaction in some individuals. However, reactions tend to be mild, and are the result of an allergic reaction. To treat a mild allergic reaction to urticating hairs: rinse the affected area with cool water, then apply a cold compress or calamine lotion to reduce itching and inflammation. Over-the-counter antihistamines may also be helpful in relieving symptoms. If the reaction is severe or if symptoms persist, it is important to seek medical attention.